Life Insecta Lepidoptera Ditrysia Papilionoidea Nymphalidae Heliconiinae

700x504 (~51Kb) USA: Corral Pass region (47°00'15N 121°29'02W), Pierce Co., WA, 18.7.2000, Photo © Markku Savela

The exact identification of this species is still unknown, but tentatively assumed to belong into this group.

• Boloria pales (Denis & Schiffermüller, 1775) Cantabrian, Pyrenees, Alps, Carpathians, Caucasus, C.Asia, W.China. See [maps] Perlada alpina Shepherd's Fritillary Dostojka pales

  • Papilio pales Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 177
  • Papilio arsilache Esper, 1780; Die Schmett. Th. I, Bd. 2 (1): 35, pl. 56, f. 5
  • Argynnis pales v. caucasica Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 9
  • Argynnis pales var. inducta Sandberg, 1883; Ent. Tidskr. 4 (3): 129, 130, (4):224
  • Argynnis pales ab. killiasi Rühl, 1892; Soc. Ent. 7 (15): 113; TL: Rhaetia
  • Argynnis pales ab. nigra Aigner-Abafi, 1906; Ann. hist.-nat. Mus. Hung. 4 (2): 502
  • Argynnis pales ab. boxhoi Lambillion, 1906; Rev. Mens. Soc. ent. namur. 6 (11): 52
  • Argynnis pales darjana Seitz, 1909; Gross-Schmett. Erde 1: 230, pl. 68 a (Stgr. i.l.); TL: Syr Darja
    • = ?Clossiana hegemone hegemone; [BRU2], 46
  • Argynnis pales ab. conducta Schultz, 1908; Ent. Zs. 22 (10): 39*; TL: Preda
  • Argynnis pales ab. deflavata Wagner, 1909; Ent. Zs. 23 (4): 17
  • Brenthis pales arsilache f. ♀ berolinensis Hannemann, 1915; Int. Ent. Zs. 9 (21): 115
  • Argynnis pales f. radiata Nordström, 1918; Ent. Tidskr. 39: 312, f. 2
  • interligata (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 46
  • hockacensis (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 51
  • othello (Cornelsen, 1923) (Argynnis); Int. Ent. Zs. 16: 212
  • elongata (Le Charles, 1926) (Argynnis); Amat. Papillons 3: 120; TL: Savoy
  • zehlae (Bois-Reymond, 1926) (Argynnis); Int. ent. Zs. 20: 105
  • alba (Cabeau, 1930) (Argynnis); Lambillionea 30: 178
  • argentaea (Mairlot, 1932) (Argynnis); Lambillionea 32: 209; TL: Belgia
  • Argynnis pales contempta Zerny, 1932; Denkschr. Akad. Wiss. Wien 103: 70
  • Argynnis pales majellensis Dannehl, 1933; Ent. Zs. 46 (23): 244; TL: Montagna Grande
  • parvula (Collier, 1933) (Argynnis); Ent. Rundsch. 50: 55
  • carelica (Valle, 1935) (Brenthis); Anim. fenn. 2 I Diurna: 80; TL: Finland
  • Argynnis pales palinoida Seok, 1937; Annot. zool. jap. 16 (2): 109; TL: Mt Sioutaisan
  • mixta Warren, 1944; Trans. R. Ent. Soc. Lond. 94 (1): 59
  • nitida Warren, 1944; Trans. R. Ent. Soc. Lond. 94 (1): 62
  • subalpina (Petersen, 1947) (Argynnis); Zool. Bidr. Uppsala 26: 404
  • scandinavica (Petersen, 1947) (Argynnis); Zool. Bidr. Uppsala 26: 406
  • Boloria pales f. maxima Verity, 1950; Le Farfalle diurn. d'Italia 4: 217
  • Boloria pales palustris f. posticeporrecta Verity, 1950; Le Farfalle diurn. d'Italia 4: 217
  • Boloria pales palustris sr. fuscata Verity, 1950; Le Farfalle diurn. d'Italia 4: 222
  • Argynnis pales var. arsilache ab. marginata Schnaider, 1950; Polskie Pismo. Ent. 19 (3-4): 242
  • pontica de Lesse & Crosson du Cormier, 1957; Lambillionea 57: 35
  • Brenthis pales rufina Agenjo, 1962; Eos 38 (3): 337; TL: Ull de Ter, 2391m, Setcasas, Gerona, Spain
  • tatrensis Crosson du Cormier, 1961; Acta. faun. ent. Mus. nat. Pragae 7: 58
  • carpathomeridionalis Crosson du Cormier & Popescu-Gorj, 1963; Acta. faun. ent. Mus. nat. Pragae 9: 145
  • ignita Krzywicki, 1963; Ann. Zool. 21: 207, 218; TL: Tatra Mts

  • Argynnis pales ab. inducta; Sahlberg, 1891, Medd. Soc. Fenn. 18: f. 6
  • Boloria pales carpathomeridionalis; [LepIndex]; [CoL]
  • Boloria pales caucasica; [LepIndex]; [CoL]
  • Boloria pales contempta; [LepIndex]; [CoL]
  • Boloria pales darjana; [LepIndex]; [CoL]
  • Boloria pales fuscata; [LepIndex]; [CoL]
  • Boloria pales ignita; [LepIndex]; [CoL]
  • Boloria pales inducta; [LepIndex]; [LepIndex]; [CoL]
  • Boloria pales killiasi; [LepIndex]; [CoL]
  • Boloria pales maxima; [LepIndex]; [CoL]
  • Boloria pales nitida; [LepIndex]; [CoL]
  • Boloria pales palinoida; [LepIndex]; [CoL]
  • Boloria pales parvula; [LepIndex]; [CoL]
  • Boloria pales pontica; [LepIndex]; [CoL]
  • Boloria pales posticeporecta; [LepIndex]; [CoL]
  • Boloria pales radiata; [LepIndex]; [CoL]
  • Boloria pales rufina; [LepIndex]; [CoL]
  • Boloria pales scandinavica; [LepIndex]; [CoL]
  • Boloria pales subalpina; [LepIndex]; [CoL]
  • Boloria pales tatrensis; [LepIndex]; [CoL]
  • Boloria pales; [H&R], 98; [BOW]: pl. 1, f. 17, 23; [EBW]; [MRS], 478; [Otakar Kudrna]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SZMN] ; [Mario Maier]
  • Boloria selene pales; [LepIndex]; [CoL]
  • Brenthis pales; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • ?Papilio arsilache; Knoch, 1781, Beitr. Insektengesch. 1: 73

  Misidentified/misapplied:

  • Papilio pales; Bergsträsser, 1780, Nomen. Ins. 4: 31, pl. 84, f. 6-7

  Larva on Viola , V. calcarata [H&R]

  • B. p. pales Austria

    • Boloria pales pales; [H&R], 98; [iNaturalist]

    Larva on Viola calcarata [H&R]

  • B. p. sifanica (Grum-Grshimailo, 1891) Qinghai, Gansu

    • Argynnis pales var. sifanica Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 456
    • Boloria pales mingi Kocman, 1995
      • = ; Huang, 2001
    • ?pales lucida (Bang-Haas, 1927) (Argynnis); Horae Macrolep. Palaearct. 1: 52
      • = ; Huang, 2001

    • Boloria pales lucida; [LepIndex]; [CoL]
    • Boloria pales sifanica; Huang, 2001
    • Boloria sifanica; [EBW], (name only); [LepIndex] ; [CoL] ; [iNaturalist]
  • B. p. eupales (Fruhstorfer, 1903) S.Tibet

    • Argynnis pales eupales Fruhstorfer, 1903; Soc. Ent. 18 (16): 124; TL: Khamba-Jong, nr Yadong of SC.Tibet
    • Argynnis pales eupales Fruhstorfer, 1904; Dt. ent. Z. Iris 16 (2): 307; TL: Khamba-Jong, S.Tibet

    • Boloria pales eupales; Huang, 2001; [LepIndex]; [CoL] ; [iNaturalist]
  • B. p. palina (Fruhstorfer, 1904) W.Sichuan

    • Argynnis pales palina Fruhstorfer, 1904; Dt. ent. Z. Iris 16 (2): 306; TL: Ts-Tsien-Lou; Siao-Lou, Setchuan
    • Boloria pales palinoides Reuss, 1925; Int. ent. Zs. 19: 280
      • = ; Huang, 2001
    • Boloria pales f.geogr. palinoides Draeseke & Reuss, 1925; Dt. ent. Z. Iris 39: 218
    • Argynnis pales palina f. conjuncta Eisner, 1942; Zool. Meded. Leiden 24 (4): 124; TL: Szetschwan
    • Argynnis pales palina f. deficiens Eisner, 1942; Zool. Meded. Leiden 24 (4): 124; TL: Szetschwan

    • Boloria pales palina; [MRS], 478; Huang, 2001
    • Boloria palina; [iNaturalist]
    • Boloria sifanica palina; [LepIndex]; [CoL]
    • Boloria sifanica palinoides; [LepIndex]; [CoL]
  • B. p. palustris (Fruhstorfer, 1909) S.France (Alpes Maritimes), Switzerland (Alps)

    • Argynnis pales palustris Fruhstorfer, 1909; Int. ent. Zs. 3 (20): 112; TL: Simplon-Culm, Zermatt, Arolla, Engadin

    • Boloria pales palustris; [H&R], 99
  • B. p. pyrenesmiscens Verity, 1932 Pyrenees

    • Boloria pales r. pyrenesmiscens Verity, 1932; Dt. ent. Z. Iris 46 (3): 102; TL: Pyrenees

    • Boloria pales pyrenesmiscens; [H&R], 99; [iNaturalist]
    • Boloria pales pyrenesniscens; [LepIndex]; [CoL]
    • Boloria pales pyrenesniscens[sic, pyrenesmiscens]; Warren, 1944, Trans. R. Ent. Soc. Lond. 94 (1): 79
  • B. p. rilaensis Varga, 1971 Bulgaria

    • Boloria pales rilaensis Varga, 1971; Acta. Biol. Debrecina (2) 9: 215; TL: Bulgaria, Rila Planina, Malak Mecsi

    • Boloria pales rilaensis; [SZMN]
    • Boloria pales rilaoensis; [LepIndex]; [CoL]
  • B. p. yangi Hsu & Yen, 1997 Taiwan

    • Boloria pales yangi Hsu & Yen, 1997; J. Res. Lep. 34: 143; TL: Taiwan, Lishan (24°15N, 121°14E)
  • B. p. shambhala Kocman, 1999

    • Boloria pales shambhala Kocman, 1999; ; TL: SE.Qinghai
  • B. p. nirvana Kocman, 1999

    • Boloria pales nirvana Kocman, 1999; ; TL: Lhasa
• Boloria sipora (Moore, [1875]) -. See [maps]

  • Argynnis sipora Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4): 568, pl. 66, f. 11; TL: Kashmere, nar Gungabul lake, at foot of Haramook Peak

  • Boloria pales sipora; [LepIndex]; [CoL]
  • Boloria sipora; [BRU2]: 53, pl. 35, f. 12-18; [iNaturalist] ; [LEPIFORUM]
  • B. s. sipora E.Pamirs

    • Argynnis baralacha Moore, 1882; Proc. zool. Soc. Lond. 1882 (1): 242, pl. 11, f. 1, 1a; TL: Ladak
    • Argynnis pales hunzaica Tytler, 1940; J. Bombay nat. Hist. Soc. 42 (1): 120; TL: Mizgah, Hunza

    • Argynnis baralacha; [LepIndex] ; [CoL]
    • Boloria pales baralacha; [LepIndex]; [CoL]
    • Boloria sipora sipora; [BRU2], 53; [iNaturalist]
  • B. s. korla (Fruhstorfer, 1904)

    • Argynnis pales korla Fruhstorfer, 1904; Dt. ent. Z. Iris 16 (2): 306; TL: Korla

    • Boloria pales korla; [LepIndex]; [CoL]

    800x600 (~99Kb) underside An ancient moraine at 3400 m above sea level in the Tyshkan River headwaters, 45 km NEN of Zharkent, 20 km NEN of village Sarybel', altitude of 3400 m, mountain massif Tyshkantau, the Southern Dzhungarian Alatau mountain chain (NE part of Tien Shan), Zharkent District, Taldy-Kurgan Province, SE Kazakhstan. 26th of July 1994, Photo © Oleg Kosterin

• Boloria generator (Staudinger, 1886) W.Himalaya. See [maps]

  • Argynnis pales var. generator Staudinger, 1886; Stett. Ent. Ztg. 47 (7-9): 235; TL: Alai Mts

  • Argynnis pales var. generator; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 438, pl. 12, f. 4a-b
  • Boloria generator; [Oleg Kosterin]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34; [LEPIFORUM]
  • B. g. generator Altai

    • Argynnis pales var. juldussica Wagner, 1913; Ent. Mitt. 2 (4): 115; TL: Juldus
    • Argynnis (Brentis[sic]) pales generator ab. extrema Wnukowsky, 1927; Mitt. Münch. Ent. Ges. 17 (7-12): 71; TL: Altai

    • Boloria generator generator; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Boloria pales generator; [LepIndex]; [CoL]
    • Boloria sipora generator; [BRU2], 53; [iNaturalist]
  • B. g. klimenkoi Churkin & Zhdanko, 2001 Tien Shan

    • generator klimenkoi Churkin & Zhdanko, 2001; Tethys Ent. Res. 3: 153; TL: Kyrghyzstan, W. Tien-Shan, Chatkal Mts, Sary-Chelek

    • Boloria generator klimenkoi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
• Boloria napaea (Hoffmannsegg, 1804) Pyrenees, Alps, Fennoscandia. See [maps] Napaea Fritillary Tunturihopeatäplä Nordisk Gullfläcklig Pärlemorfjäril Fjällpärlefjäril Mountain Fritillary

  • Papilio napaea Hoffmannsegg, 1804; Mag. f. Insektenk. 3: 196 (repl. Papilio isis Hübner, [1799-1800])
  • Papilio isis Hübner, [1799-1800]; Samml. eur. Schmett. [1]: 9, pl. 7, f. 39-38 (preocc. Drury, 1773)
  • Papilio dirphya Hoffmannsegg, 1806; Mag. f. Insektenk. 5: 180 (repl. Papilio napaea Hoffmannsegg, 1804)
  • ?Papilio napaea Hübner, [1819]; Samml. eur. Schmett. [1]: pl. 151, f. 757-758
  • Boloria pales f. atroviolacea Verity, 1932; Dt. ent. Z. Iris 46 (3): 108
  • Boloria pales f. atroviridans Verity, 1932; Dt. ent. Z. Iris 46 (3): 108
  • frigida Warren, 1944; Trans. R. Ent. Soc. Lond. 94 (1): 46; TL: N.Scandinavia
  • Boloria napaea napaea f. infrapallida Verity, 1950; Le Farfalle diurn. d'Italia 4: 215
  • Boloria napaea napaea f. laterubra Verity, 1950; Le Farfalle diurn. d'Italia 4: 215
  • Boloria napaea f. latevirida Verity, 1950; Le Farfalle diurn. d'Italia 4: 215
  • Boloria napaea napaea f. nitidarubra Verity, 1950; Le Farfalle diurn. d'Italia 4: 215
  • Boloria napaea napaea sr. modesta Verity, 1950; Le Farfalle diurn. d'Italia 4: 216
  • pyreneorientalis de Lesse, 1962; Alexanor 1: 232; TL: France
  • dolomitensis Crosson du Cormier & Guerin, 1964; Alexanor 3: 313; TL: Italy
  • inalpina Crosson du Cormier & Guerin, 1964; Alexanor 3: 313; TL: Switzerland
  • pennina Crosson du Cormier & Guerin, 1964; Alexanor 3: 314; TL: Italy
  • sabaudiensis Crosson du Cormier & Guerin, 1964; Alexanor 3: 314
  • punctata Crosson du Cormier & Guerin, 1964; Alexanor 3: 315
  • laurentidis Crosson du Cormier, 1966; Alexanor 4: 291

  • Argynnis napaea; Hübner, [1819], Verz. bek. Schmett. (2): 30
  • Boloria napaea dolomitensis; [LepIndex]; [CoL]
  • Boloria napaea frigida; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
  • Boloria napaea inalpina; [LepIndex]; [CoL]
  • Boloria napaea infrapallida; [LepIndex]; [CoL]
  • Boloria napaea laterubra; [LepIndex]; [CoL]
  • Boloria napaea latevirida; [LepIndex]; [CoL]
  • Boloria napaea laurentidis; [LepIndex]; [CoL]
  • Boloria napaea modesta; [LepIndex]; [CoL]
  • Boloria napaea nitidarubra; [LepIndex]; [CoL]
  • Boloria napaea pennina; [LepIndex]; [CoL]
  • Boloria napaea punctata; [LepIndex]; [CoL]
  • Boloria napaea pyreneorientalis; [LepIndex]; [CoL]
  • Boloria napaea sabaudiensis; [LepIndex]; [CoL]
  • Boloria napaea; [SPRK]; [H&R], 99; [MRS], 478; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SZMN] ; [SVF] ; [Mario Maier]

  Misidentified/misapplied:

  • Boloria napaea (= Boloria alaskensis); [NACL], #4462
  • Boloria napaea (= Boloria alaskensis); [BNA]: 317, #237, pl. 26
  • Boloria napaea (= Boloria alaskensis); [NAB], 557

  Larva on Polygonum [SPRK], Polygonum viviparum [NAB]

  • B. n. altaica (Grum-Grshimailo, 1893) Tarbagatai, Sayan, Altai - S.Transbaikalia, Mongolia

    • Argynnis pales var. altaica Grum-Grshimailo, 1893; Horae Soc. ent. Ross. 27 (1-2): 128; TL: [Kara-Kaba Valley, Altai]
    • Argynnis arsilache isisoides Heydemann, 1920; Dt. Ent. Z. Iris 34: 271

    • Boloria altaica altaica; [BRU2], 53
    • Boloria altaica; [BRU2]: 53, pl. 35, f. 19-24; Yakovlev, 2012, Nota lepid. 35 (1): 91; [iNaturalist]
    • Boloria napaea altaica; [MRS], 478; [Oleg Kosterin]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34; [LepIndex]; [CoL]

    400x533 (~48Kb) ♂ underside An alpine meadow, 2200 m above sea level, on the eastern spurs of the Katunskiy Mt. Range, the valley of the Argem (Direntai) stream, Central Altai Mts., West Siberia, Russia. 17th July, 1988 (on an Aster alpinum L. flower), Photo © Oleg Kosterin	652x500 (~54Kb) ♂ ♀ upperside A damp meadow on the Samakha Rivulet bank in the hollow Step' Samakha, 1600 m above sea level, at the junction of Argut and Koksu Rivers, Kosh-Agach District, Altai Republic, Central Altai Mts., West Siberia, Russia. 9th July, 1988 (on Bistorta bistorta), Photo © Oleg Kosterin	538x464 (~27Kb) ♂ ♀ upperside underside Highland dwarf birch (Betula rotundifolia) tundras on a ledge of the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2400 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 10th July 1998, Photo © Oleg Kosterin
    790x560 (~86Kb) ♂ upperside a subalpine parkland of Pinus sibirica Du Tour on the Seminskii Pass through the Seminskii Mt. Range, at 1800 m above sea level. Ongudai District, Altai Republic, West Siberia, Russia. 9th July 1999 (on Betula nana rotundifolia), Photo © Oleg Kosterin	498x578 (~68Kb) ♂ underside a subalpine parkland of Pinus sibirica Du Tour on the Seminskii Pass through the Seminskii Mt. Range, at 1800 m above sea level. Ongudai District, Altai Republic, West Siberia, Russia. 9th July 1999 (on Aster alpinus), Photo © Oleg Kosterin

  • B. n. shanxiana Murayama, 1983

    • Boloria napaea shanxiana Murayama, 1983; Entomotaxonomia 5: (281-288)
  • B. n. vinokurovi Dubatolov, 1992 E.Yakutia

    • Boloria napaea vinokurovi Dubatolov, 1992; Vestn. Zool. 6: 40

    • Boloria altaica vinokurovi; [BRU2], 53
    • Boloria napaea vinokurovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34; [SZMN]
  • B. n. pustagi Korshunov & Ivonin, 1995 NE.Altai

    • Boloria altaica pustagi Korshunov & Ivonin, 1995; ; TL: Gornaya Shoriya, NE. Altai
      • = Boloria napaea altaica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34

    • Boloria altaica pustagi; [BRU2], 53
    • Boloria napaea pustagi; [SZMN]
  • B. n. contaminata Kosterin & Gorbunov, 2007 N.Ural

    • Boloria napaea contaminata Kosterin & Gorbunov, 2007; Butt. North. Asia in nat. 2: 186

    • Boloria napaea contaminata; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
• Boloria purpurea Churkin, 1999 Buryatia (Barguzin Mts.). See [maps]

  • Boloria purpurea Churkin, 1999; Atalanta 29 (1/4): 118, pl. IX, f. 1-3; TL: Buryatia, Barguzin Mts., Nesterikha River, Kedrovoe Lake, 1900-2000m

  • Boloria frigidalis purpurea;
  • Boloria purpurea; [BRU2]: 54, pl. 36, f. 10-12; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
• Boloria alaskensis (Holland, 1900) Alaska - British Columbia, Arctic America. See [maps]

  • Brenthis pales var. alaskensis Holland, 1900; Ent. News 11 (3): 383; TL: mtns betw. Forty-Mile and Mission Creeks [Alaska]

  • Boloria (Boloria) alaskensis alaskensis; [P&N23]
  • Boloria (Boloria) alaskensis; Pelham, 2008, J. Res. Lepid. 40: 296
  • Boloria alaskensis; [BRU2]: 54, pl. 36, f. 1-9; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [RFEL]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BAMONA]
  • Boloria napaea alaskensis; [NACL], #4462a; [BNA], #237
  • Brenthis pales alaskensis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16

  Larva on Polygonum viviparum [BNA]: 317, Polygonum , Viola [BRU2]: 55

  • B. a. alaskensis E.Chukotka, Wrangel I., Alaska - Alberta

    • Boloria reiffi Reuss, 1925; Int. ent. Zs. 19: 279; TL: Br. Columbia
      • = ; Pelham, 2008, J. Res. Lepid. 40: 296
    • Boloria pales arsilache r. nearctica Verity, 1932; Dt. ent. Z. Iris 46 (3): 104
      • = ; Pelham, 2008, J. Res. Lepid. 40: 296

    • Boloria alaskensis alaskensis; [BRU2], 54; Pelham, 2008, J. Res. Lepid. 40: 296; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [RFEL]; [iNaturalist] ; [BOA]
    • Boloria napaea nearctica; [NACL], #4462b; [LepIndex]; [CoL]
    • Boloria napaea reiffi; [LepIndex]; [CoL]
    • Boloria pales nearctica; [LepIndex]; [CoL]
  • B. a. nikolajewski (Heydemann, 1920) Amur

    • Argynnis arsilache nikolajewski Heydemann, 1920; Dt. Ent. Z. Iris 34: 271 (Bang-Haas i.l.); TL: Amur
      • = ?Boloria alaskensis; [BRU2], 54 (?)

    • Boloria alaskensis nikolajewski; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [RFEL]
    • Boloria napaea nikolajewski; [LepIndex]; [CoL]
  • B. a. halli Klots, 1940 Wyoming

    • Boloria pales halli Klots, 1940; Amer. Mus. Novit. 1054: 1; TL: vic. Green River Pass, Wind River Range, Sublette Co., Wyoming

    • Boloria (Boloria) alaskensis halli; [P&N23]
    • Boloria alaskensis halli; Pelham, 2008, J. Res. Lepid. 40: 296; [iNaturalist] ; [BOA]
    • Boloria napaea halli; [NACL], #4462c; [BNA], #237; [LepIndex]; [CoL]

    Larva on Polygonum bistortoides? [BNA]: 317

  • B. a. sedykhi Crosson du Cormier, 1977 Polar Urals, Yamal Peninsula

    • Boloria alaskensis sedykhi Crosson du Cormier, 1977; Alexanor 10 (1): 46

    • Boloria alaskensis sedykhi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [LepIndex]; [CoL]
  • B. a. bato Churkin, 1999 Transbaikalia

    • Boloria alaskensis bato Churkin, 1999; Atalanta 29 (1/4): 117; TL: Kodar Mts., Chita Region

    • Boloria alaskensis bato; [BRU2], 54; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35
• Boloria frigidalis Forster, 1968 Altai, Sayan, Tuva (mountains), W.Mongolia. See [maps]

  • frigidalis Forster, 1968; Reichenbachia 11 (18): 189; TL: ;Tchuja Mts., SE Altai
  • Boloria napaea altaica f. frigidalis Warren, 1944; Trans. R. Ent. Soc. Lond. 94 (1): 48; TL: Tchuja Mts., SE Altai

  • Boloria frigidalis; Forster, 1968, Reichenbachia 11 (18): 189-205; Kosterin, 1994, Actias. Russian Journal of Scientific Lepidopterology,1 (1-2): 46-76; [BRU2]: 54, pl. 35, f. 25-27; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; Yakovlev, 2012, Nota lepid. 35 (1): 91; [iNaturalist] ; [LEPIFORUM]

  300x496 (~17Kb) ♀ underside An alpine meadow vegetation at a brook flowing in a moss/sedge tundra in a cirque of the sources of a rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2650 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16 July 1998 (on Rhodiola algida (Ledeb.) Fisch. & Mey.), Photo © Oleg Kosterin	960x588 (~107Kb) ♀ underside A chionophylic alpine meadow in a cirque of one of the Chikty Rivulet sources, 2700 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 13th July 1998 (on Allium schoenoprasum), Photo © Oleg Kosterin	482x654 (~23Kb) ♂ underside A chionophylic alpine meadow in a cirque of one of the Chikty Rivulet sources, 2700 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 13th July 1998 (on Allium schoenoprasum), Photo © Oleg Kosterin
  864x712 (~99Kb) ♂ underside A chionophylic alpine meadow in a cirque of one of the Chikty Rivulet sources, 2700 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 13th July 1998 (on Allium schoenoprasum), Photo © Oleg Kosterin	597x400 (~38Kb) ♂ upperside A chionophylic alpine meadow in a cirque of one of the Chikty Rivulet sources, 2700 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 13th July 1998, Photo © Oleg Kosterin

• Boloria caucasica (Lederer, 1852) Caucasus, Transcaucasia, Turkey. See [maps]

  • Argynnis arsilache v. caucasica Lederer, 1852; Verh. zool.-bot. Ver. Wien 2 (Abh.): 22 (Bisch. i.l.); TL: "Achaltsiche und Kazbegi" [Akhaltsike and Kazbegi, Georgia]
  • Boloria pontica Crosson du Cormier, Guérin & de Lesse, 1957

  • Boloria caucasica; [BRU2]: 52, pl. 35, f. 28-30; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34; [iNaturalist] ; [LEPIFORUM]

  1016x908 (~323Kb) upperside Russia, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, Blyam Mt. E. slope, 12 July 2008, Photo © Oleg Kosterin

  1242x874 (~308Kb) underside Russia, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, Blyam Mt. E. slope, 12 July 2008, Photo © Oleg Kosterin

  908x851 (~177Kb) Russia, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, Blyam Mt. E. slope, 12 July 2008, Photo © Oleg Kosterin

  • B. c. causasica Caucasus, Transcaucasia, Turkey
  • B. c. petrovi Churkin, 2002 Daghestan

    • Boloria caucasica petrovi Churkin, 2002; Helios 3: 109, pl. 8, f. 10-13; TL: Daghestan
      • = Boloria caucasica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
• Boloria aquilonaris (Stichel, 1908) W.Siberia, C.Siberia. See [maps] Tranpärlefjäril Suohopeatäplä Dostojka akwilonaris Allmän gulfläckig pärlemofjäril Cranberry Fritillary

  • Argynnis pales aquilonaris Stichel, 1908; Berl. ent. Z. 53 (1): 81, pl. 3, f. 5; TL: Gallivar, Sweden
  • parvula (Collier, 1933) (Argynnis); Ent. Rundsch. 50: 55; TL: Germany
  • ericiti Crosson du Cormier & Guerin, 1959; Alexanor 1: 66; TL: Switzerland
  • toulgoeti Crosson du Cormier & Guerin, 1959; Alexanor 1: 67; TL: Switzerland
  • podhalensis Crosson du Cormier & Guerin, 1962; Alexanor 2: 237; TL: Poland

  • Boloria aquilonaris ericiti; [LepIndex]; [CoL]
  • Boloria aquilonaris podhalensis; [LepIndex]; [CoL]
  • Boloria aquilonaris toulgueti; [LepIndex]; [CoL]
  • Boloria aquilonaris; [H&R], 102; [BRU2]: 52, pl. 36, f. 13-18; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SVF] ; [Mario Maier]

  Larva on Vaccinium , Oxycoccus palustris, Oxycoccus microcarpus [BRU2]: 52

  • B. a. aquilonaris Fennoscandia, N.European Russia

    • Argynnis pales var.[?] lapponica Staudinger, 1861; Stettin Ent. Ztg 22 (10-12): 347
    • Argynnis pales lapponica Staudinger, 1871; (preocc. Esper)
    • Brenthis pales arsilache f. hannoverana Füge, 1912; Zs. wiss. Insektenbiol. 8 (5): 189
    • Boloria pales hypoarsilache Verity, 1932; Dt. ent. Z. Iris 46 (3): 106 (repl. lapponica Staudinger)
    • alethea (Hemming, 1934) (Argynnis); Stylops 3 (5): 97
    • Boloria alethea nigrofasciata Krzywicki, 1967; Ann. zool. Warsz. 25 (1): 115

    • Boloria alethea hannoverana; [LepIndex]; [CoL]
    • Boloria alethea nigrofasciata; [LepIndex]; [CoL]
    • Boloria alethea; [LepIndex] ; [CoL]
    • Boloria aquilonaris aquilonaris; [BRU2], 52; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [SZMN]
  • B. a. sima Churkin, 2000 Kazakhstan

    • Boloria aquilonaris sima Churkin, 2000; Helios 1: 107
      • = Boloria aquilonaris aquilonaris; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35

    • Boloria aquilonaris sima; [BRU2], 52
  • B. a. huerteri Churkin & Bogdanov, 2003 Buryatia

    • Boloria aquilonaris huerteri Churkin & Bogdanov, 2003; Helios 4: 217; TL: Russia, Siberia, Buryatia, Dawan Pass, 1400m

    • Boloria aquilonaris huerteri; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35
• Boloria banghaasi Seitz, 1909 Amur, E.Yakutia?, Kamchatka, Sakhalin?. See [maps]

  • Boloria pales f. banghaasi Seitz, 1909; Gross-Schmett. Erde 1: 231, pl. 68 a; TL: "Kentei" [Kentei Mts., ?Mongolia]

  • Boloria banghaasi; [BRU2]: 52, pl. 35, f. 31-36; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [RFEL]; [iNaturalist]

  533x400 (~50Kb) underside On the local dwarf birch Betula divaricata Ledeb. An Empetrum/lichen tundra alternating with dwarf pine Pinus pumila (Pall.) Regel elfin wood in the delta of the Burgauli River, the southern coast of the Koni Peninsula, Ola District, Magadan Province, Russia. 18th July, 1989, Photo © Oleg Kosterin

  • B. b. banghaasi

    • Boloria aquilonaris banghaasi; [Oleg Kosterin]
    • Boloria banghaasi banghaasi; [BRU2], 53; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35
    • Boloria sifanica banghaasi; [LepIndex]; [CoL]
  • B. b. neopales (Nakahara, 1926) Sakhalin

    • Argynnis pales neopales Nakahara, 1926; Ins. Inscit. Mens. 14: 48 (repl. Argynnis pales sachalinensis Matsumura, 1925); TL: Sakhalin
      • = ?Boloria banghaasi; [BRU2], 53 (?)
    • Argynnis pales sachalinensis Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15: 100 (preocc.); TL: Saghalin

    • Boloria banghaasi neopales; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [RFEL]
    • Boloria pales neopales; [LepIndex]; [CoL]
    • Boloria selene sachalinensis; [CoL]
  • B. b. infans Churkin, 2000 Magadan, Chukotka

    • Boloria banghaasi infans Churkin, 2000; Helios 1: 109; TL: Chukotka, Bilibino

    • Boloria banghaasi infans; [BRU2], 53; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [RFEL]
  • B. b. jakubovi Kosterin & Gorbunov, 2007 S.Kamtchatka

    • Boloria aquilonaris jakubovi Kosterin & Gorbunov, 2007; Butt. North. Asia in nat. 2: 197; TL: S.Kamchatka, 52°25N, 156°26E

    • Boloria banghaasi jakubovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; [RFEL]
• Boloria roddi Kosterin, 2000 Altai. See [maps]

  • Boloria pales roddi Kosterin, 2000; Far East Ent. 86 (1): 3; TL: Alatai, Chuiskaya Step, Kosh-Agach
  • sanguinea Churkin & Bogdanov, 2002
    • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35

  • Boloria banghaasi roddi; [BRU2], 53
  • Boloria roddi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 35; Yakovlev, 2012, Nota lepid. 35 (1): 91; [iNaturalist]

  598x526 (~60Kb) ♂ upperside Boggy, with Pentaphylloides fruticosa bushes left bank of the Dzhazator River in its upper flow, between the falls of its right tributaries Chikty and Akbul, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 19th July 1998, Photo © Oleg Kosterin

  732x500 (~79Kb) ♂ upperside Boggy, with Pentaphylloides fruticosa bushes left bank of the Dzhazator River in its upper flow, between the falls of its right tributaries Chikty and Akbul, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 19th July 1998, Photo © Oleg Kosterin

  400x515 (~34Kb) ♂ underside Boggy, with Pentaphylloides fruticosa bushes left bank of the Dzhazator River in its upper flow, between the falls of its right tributaries Chikty and Akbul, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 19th July 1998 (on Tripleurospermum ambiguum), Photo © Oleg Kosterin

• Boloria graeca (Staudinger, 1870) Balkans, Alps. See [maps] Balkan Fritillary

  • Argynnis pales var. graeca Staudinger, 1870; Horae Soc. ent. ross. 7 (1870): 62, pl. 1, f. 4
  • higginsi Praviel, 1938; Rev. fr. Lépid 9: 168

  • Boloria graeca higginsi; [LepIndex]; [CoL]
  • Boloria graeca; [H&R], 103; [Otakar Kudrna]; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE]
  • B. g. graeca

    • Boloria graeca graeca; [H&R], 103
  • B. g. balcanica (Rebel, 1903) Bulgaria

    • Argynnis pales var. balcanica Rebel, 1903; Ann. nat. Mus. Wien 18: 169
    • Boloria (Smoljana) rhodopensis Slivov, 1995; Acta zool. bulg. 48: 63
      • = ; Abadjiev & Beshkov, 2000, Phegea 28 (1): 24

    • Boloria graeca balcanica; [H&R], 103; [LepIndex]; [CoL]
  • B. g. brogotarus (Fruhstorfer, 1909) Alpes Maritimes

    • Argynnis pales brogotarus Fruhstorfer, 1909; Int. ent. Zs. 3 (20): 112; TL: Alpes Maritimes
    • medioitalica (Turati, 1912) (Argynnis); Ann. Mus. Napoli 3 (18): 21
    • tendensis (Higgins, 1930) (Argynnis); Entomologist 63: 202

    • Boloria graeca brogotarus; [LepIndex]; [CoL]
  • B. g. karina van Oorschot & Wagener, 1990 Turkey

    • Boloria graeca karina van Oorschot & Wagener, 1990; Ent. Ber. 50 (9): 117; TL: Turkey, Erzurum, Palandöken Daglan, 5 km S Erzurum, 2200-2500m

• Boloria eunomia (Esper, 1800) . See [maps] Bog Fritillary Randring-Perlmutterfalter Rämehopeatäplä Svarttringlad pärlemorfjäril Dostojka eunomia Odonpärlefjäril Ocellate Bog Fritillary

  • Papilio eunomia Esper, 1800; Die Schmett., Suppl. Th 1 (8-9): 94, pl. 110, f. 5; TL: Koenigberg
  • Argynnis aphirape ab. haverkampfi Ball, 1912; Ann. Soc. ent. Belg. 56: 280
  • Boloria aphirape helmina Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 17; TL: E.Preuss, Zehlau-Bruch
  • Argynnis aphirape ceretanensis Deslandes, 1930; Bull. Soc. ent. Fr. 35 (15): 243; TL: Porté (Pyr.-Orientales), 1600m
  • Argynnis aphirape ceretanensis f. sublatefasciata Deslandes, 1930; Bull. Soc. ent. Fr. 35 (15): 244
  • melanira (Cabeau, 1930) (Argynnis); Lambillionea 30: 178; TL: Belgium
  • diluta (Heydemann, 1942) (Argynnis); Dt. ent. Z. Iris 55: 94
  • subargentata (Petersen, 1947) (Argynnis); Zool. Bidr. Uppsala 26: 389
  • montana (Petersen, 1947) (Argynnis); Zool. Bidr. Uppsala 26: 398
  • Proclossiana eunomia f. suprapupillata Verity, 1950; Le Farfalle diurn. d'Italia 4: 226
  • Proclossiana eunomia gieysztori Krzywicki, 1967; Ann. zool. Warsz. 25 (1): 116

  • Boloria (Boloria) eunomia; Pelham, 2008, J. Res. Lepid. 40: 296
  • Boloria (Proclossiana) eunomia; [FE]; [P&N23]
  • Boloria eunomia; [Opler]; [BNA]: 318, #238, pl. 26; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [MPG] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [BOA] ; [BAMONA] ; [SVF]
  • Clossiana eunomia; [SPRK]; [BARU], #187; [RFEL]
  • Proclossiana aphirape ceretanensis; [LepIndex]; [CoL]
  • Proclossiana aphirape diluta; [LepIndex]; [CoL]
  • Proclossiana aphirape helmina; [LepIndex]; [CoL]
  • Proclossiana aphirape montana; [LepIndex]; [CoL]
  • Proclossiana aphirape subargentata; [LepIndex]; [CoL]
  • Proclossiana aphirape suprapupillata; [LepIndex]; [CoL]
  • Proclossiana eunomia gieysztori; [LepIndex]; [CoL]
  • Proclossiana eunomia haverkampfi; [LepIndex]; [CoL]
  • Proclossiana eunomia sublatefasciata; [LepIndex]; [CoL]
  • Proclossiana eunomia; [H&R], 103; [NACL], #4463; [NAB], 558; [EBW]; [MRS], 478; [BRU2]: 42, pl. 31, f. 1-12; [LepIndex] ; [CoL] ; [SZMN] ; [Mario Maier]

  450x713 (~65Kb) upperside	427x675 (~66Kb) underside	419x352 (~33Kb) upperside
  409x342 (~34Kb) underside	780x500 (~92Kb) upperside Russia, Moscow area, 15.6.2011 (36°25'E, 56°00'N), Photo © D. Smirnov	1009x577 (~114Kb) underside Russia, Moscow area, 17.6.2011 (36°25'E, 56°00'N), Photo © D. Smirnov

  Larva on Polygonum viviparum, Viola , Salix [NAB], Vaccinium uliginosum, Andromeda polifolia [SPRK]

  • B. e. eunomia CEU, SEU, N.Caucasus, W.Siberia

    • Papilio aphirape Hübner, [1799-1800]; Samml. eur. Schmett. [1]: pl. 5, f. 23-25
      • = Boloria eunomia; Pelham, 2008, J. Res. Lepid. 40: 297
    • Papilio tomyris Herbst, 1800; Naturs. Schmett. 10: 102, pl. 20, f. 6-7
      • = Boloria eunomia; Pelham, 2008, J. Res. Lepid. 40: 297
    • Argynnis aphirape ab. deanella Schmidt, 1912; Ent. Zs. 26 (2): 6
    • Argynnis aphirape ab. marga Schmidt, 1912; Ent. Zs. 26 (2): 6
    • Argynnis aphirape ab. excelsa Schmidt, 1912; Ent. Zs. 26 (2): 6
    • semiisabella (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 50
    • melanira (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 54
    • eustigmoea (Cabeau, 1924) (Argynnis); Rev. mens. Soc. ent. Namur. 24: 34
    • peleis (Cabeau, 1924) (Argynnis); Rev. mens. Soc. ent. Namur. 24: 35
    • Argynnis aphirape gen. aut. reichi Schneider, 1935; Int. ent. Zs. 29: 290
    • sarleti (Derenne, 1935) (Argynnis); Lambillionea 35: 213

    • Boloria eunomia eunomia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Proclossiana aphirape; [LepIndex] ; [CoL]
    • Proclossiana eunomia eunomia; [H&R], 104; [BRU2], 42

    Larva on Viola palustris, Trollius , Thalictrum , Gaulthera , Pentaphylloides fruticosa, Andromeda polifolia, Vaccinium uliginosum, Bistorta major, B. vivipara, Linaria vulgaris, Melampyrum [BRU2]: 43

  • B. e. acidalia (Böber, 1809) Altai, Sayan, S.Siberia, Amur, Ussuri

    • Papilio acidalia Böber, 1809; Mém. Soc. Imp. Nat. Moscou 2: 309
    • Argynnis aphirape v. asiatica Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 34
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana eunomia stromi Korshunov, 1996
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • yakovlevi Korb, 1999; Lambillionea 99 (4): [?]

    • Boloria eunomia acidalia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
    • Boloria eunomia stromi; [iNaturalist]
    • Boloria eunomia yakovlevi; [iNaturalist]
    • Clossiana eunomia acidalia; [RFEL]
    • Clossiana eunomia asiatica; Yakovlev, 2012, Nota lepid. 35 (1): 90
    • Proclossiana aphirape asiatica; [LepIndex]; [CoL]
    • Proclossiana eunomia acidalia; [Oleg Kosterin]; [BRU2], 42

    533x400 (~49Kb) ♂ underside An alpine meadow on a cirque bottom at 2500 m above sea level in the Argem (Direntai) Stream headwaters, 15 km W of the village Dzhazatr on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 19th July, 1988, Photo © Oleg Kosterin

  • B. e. ossianus (Herbst, 1800) Fennoscandia - Kamchatka, W.Chukotka, Far East

    • Papilio ossianus Herbst, 1800; Naturs. Schmett. 10: pl. 270, f. 4-5
    • Argynnis aphirape var. isabella Tengström, 1869; Notis. Sällsk. Fauna & Fl. Fenn. Förh. 10: 293
    • Argynnis aphirape ab. kullervo Sahlberg, 1891; Medd. Soc. Fenn. 18: 204, f. 1
    • Argynnis aphirape ab. aino Sahlberg, 1891; Medd. Soc. Fenn. 18: 204, f. 2
    • centripunctata (Nordström, 1935) (Brenthis); Ent. Tidskr. 56: 153; TL: Sweden
    • fasciata (Rangnow, 1935) (Argynnis); Ent. Rdsch. 52 (13): 176
    • radiata (Rangnow, 1935) (Argynnis); Ent. Rdsch. 52 (13): 176
    • Argynnis apiraphe ossianus f. fasciata Eisner, 1942; Zool. Meded. Leiden 24 (4): 122; TL: Kistrund, Finmark

    • Argynnis aphirape ab. isabella; Sahlberg, 1891, Medd. Soc. Fenn. 18: 204, f. 3
    • Argynnis aphirape var. isabella; Sahlberg, 1883, Medd. Soc. Fenn. 9: 160
    • Boloria eunomia ossianus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
    • Clossiana eunomia ossianus; [RFEL]
    • Proclossiana aphirape isabella; [LepIndex]; [CoL]
    • Proclossiana aphirape ossianus; [LepIndex]; [CoL]
    • Proclossiana eunomia ossiana; [MRS], 478; [BRU2], 43
    • Proclossiana eunomia ossianus; [H&R], 104
  • B. e. triclaris (Hübner, [1821]) Labrador - Alaska, Alberta

    • Argynnis triclaris Hübner, [1821]; Samml. exot. Schmett. 2: pl. [19]
    • Proclossiana ossianus (Boisduval, 1832); (preocc. by Herbst, 1800)
    • Argynnis lais Scudder, 1875; Proc. Boston Soc. nat. Hist. 17 (3): 296
      • = ; Pelham, 2008, J. Res. Lepid. 40: 297

    • Boloria (Proclossiana) eunomia triclaris; [P&N23]
    • Boloria eunomia triclaris; [BNA], #238; Pelham, 2008, J. Res. Lepid. 40: 297; [iNaturalist] ; [BOA]
    • Brenthis triclaris; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
    • Proclossiana aphirape triclaris; [LepIndex]; [CoL]
    • Proclossiana eunomia triclaris; [NACL], #4463a
  • B. e. dawsoni (Barnes & McDunnough, 1916) S.Manitoba - Maine

    • Brenthis aphirape dawsoni Barnes & McDunnough, 1916; Can. Ent. 48 (7): 222; TL: Hymers, Ontario
    • Brenthis aphirape dawsoni tr.f. harperi Gunder, 1934; Can. Ent. 66 (6): 127; TL: Churchill, Manitoba
      • = ; Pelham, 2008, J. Res. Lepid. 40: 297

    • Boloria (Proclossiana) eunomia dawsoni; [P&N23]
    • Boloria eunomia dawsoni; [BNA], #238; Pelham, 2008, J. Res. Lepid. 40: 297; [iNaturalist] ; [BOA]
    • Proclossiana aphirape dawsoni; [LepIndex]; [CoL]
    • Proclossiana eunomia dawsoni; [NACL], #4463c
  • B. e. nichollae (Barnes & Benjamin, 1926)

    • Brenthis aphirape r. nichollae Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 92; TL: Rock Mts

    • Boloria (Proclossiana) eunomia nichollae; [P&N23]
    • Boloria eunomia nichollae; Pelham, 2008, J. Res. Lepid. 40: 297; [iNaturalist] ; [BOA]
    • Proclossiana aphirape nichollae; [LepIndex]; [CoL]
    • Proclossiana eunomia nichollae; [NACL], #4463d
  • B. e. caelestis (Hemming, 1933) SW.Montana - Colorado

    • Argynnis aphirape caelestis Hemming, 1933; Entomologist 66 (12): 275 (repl. Brenthis aphirape alticola); TL: Hall Valley, Colorado
    • Brenthis aphirape alticola Barnes & McDunnough, 1913; Contr. nat. Hist. Lep. N. Am. 2 (3): 97 (preocc. Argynnis alticola Sushkin & Tsetverikov, 1907); TL: Hall Valley, Colorado
      • = ; Pelham, 2008, J. Res. Lepid. 40: 297

    • Boloria (Proclossiana) eunomia caelestis; [P&N23]
    • Boloria eunomia caelestis; [BNA], #238; Pelham, 2008, J. Res. Lepid. 40: 297; [iNaturalist] ; [BOA]
    • Proclossiana aphirape caelestis; [LepIndex]; [CoL]
    • Proclossiana eunomia caelestis; [NACL], #4463b
  • B. e. denali (Klots, 1940) Alaska

    • Brenthis (Clossiana) aphirape denali Klots, 1940; J. N.Y. Ent. Soc. 48 (4): 413; TL: McKinley National Park, Alaska

    • Boloria (Proclossiana) eunomia denali; [P&N23]
    • Boloria eunomia denali; Pelham, 2008, J. Res. Lepid. 40: 297; [iNaturalist] ; [BOA]
    • Proclossiana aphirape denali; [LepIndex]; [CoL]
    • Proclossiana eunomia denali; [NACL], #4463g
  • B. e. laddi (Klots, 1940) Wyoming

    • Clossiana aphirape laddi Klots, 1940; Amer. Mus. Novit. 1054: 4; TL: Snowy Range, Albany Co., Wyoming

    • Boloria (Proclossiana) eunomia laddi; [P&N23]
    • Boloria eunomia laddi; Pelham, 2008, J. Res. Lepid. 40: 297; [iNaturalist] ; [BOA]
    • Proclossiana aphirape laddi; [LepIndex]; [CoL]
    • Proclossiana eunomia laddi; [NACL], #4463e
  • B. e. ursadentis Ferris & Groothuis, 1971

    • Boloria eunomia ursadentis Ferris & Groothuis, 1971; J. Res. Lepid. 9 (4): 245; TL: Beartooth Pass, ½ mile S. of Montana-Wyoming border, Park Co., Wyoming

    • Boloria (Proclossiana) eunomia ursadentis; [P&N23]
    • Boloria eunomia ursadentis; Pelham, 2008, J. Res. Lepid. 40: 297; [iNaturalist] ; [BOA]
    • Proclossiana eunomia ursadentis; [NACL], #4463f; [LepIndex]; [CoL]
  • B. e. riamia (Korshunov & Ivonin, 1998) Novosibirsk

    • Clossiana[?] eunomia riamia Korshunov & Ivonin, 1998

    • Proclossiana eunomia riamina[sic, riamia]; [BRU2], 43
  • B. e. itelmena Gorbunov, 2007 Kamtchatka

    • eunomia itelmena Gorbunov, 2007; Butt. Nort. Asia in nat. 2: 183; TL: Central Kamchatka

    • Boloria eunomia itelmena; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana eunomia itelmena; [RFEL]
  • B. e. exspectata Morgun, 2011 Caucasus

    • Boloria eunomia exspectata Morgun, 2011; Caucasian Ent. Bull. 7 (1): 86; TL: Caucasus, Teberda vic., Karachaevo-Tcherkesia
  • B. e. tenera Morgun, 2011 Armenia

    • Boloria eunomia tenera Morgun, 2011; Caucasian Ent. Bull. 7 (1): 88; TL: Transcaucasia (Tsakhdadzor, Armenia)

• Boloria selene (Denis & Schiffermüller, 1775) . See [maps] Dostojka selene Ängspärlefjäril Niittyhopeatäplä Silver Meadow Fritillary Braunfleckiger Perlmutterfalter Brunfläckig pärlemorfjäril Silver-bordered Fritillary Le Petit Collier Argenté Small Pearl-bordered Fritillary Perlada castaña

  • Papilio selene Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 321; TL: Austria
  • Papilio euphrosyne Bergsträsser, 1779; Nomen. Ins. 3: 42, f. 1-2 [?]
  • Papilio euphrasia Lewin, 1795; Pap. Gr. Britain: 30, pl. 13, f. 3-4
  • Papilio marphisa Herbst, 1800; Naturs. Schmett. 10: 105, pl. 278, f. 8-9
    • = ; Pelham, 2008, J. Res. Lepid. 40: 298
  • Papilio rinaldus Herbst, 1800; Naturs. Schmett. 10: 108, pl. 271, f. 1-2, 4
    • = ; Pelham, 2008, J. Res. Lepid. 40: 298
  • Papilio silene Haworth, 1803; Lepid. Britannica (1): 34
  • Argynnis selenia Freyer, 1850; Neuere Beitr. Schmett. 6: pl. 493, f. 2
    • = ; Pelham, 2008, J. Res. Lepid. 40: 298
  • Brenthis selene var. intermedia Spångberg, 1876; Lepid. Anteckn.: 26
  • Brenthis selene ab. transversa Tutt, 1896; Brit. Butts: 295
  • Argynnis selene f. (gen. II) angustipennis Fuchs, 1899; Jb. Nassau Ver. Nat. 52: 123
  • Argynnis selene ab. flavopunctata Strand, 1900; Arch. Nat. Christian 21 (5): 20
  • Argynnis selene ab. flavopunctata Strand, 1900; Ent. Nachr. 26 (15 & 16): 226
  • Argynnis selene ab. pallida Spuler, 1901; Schmett. Eur. 1: 26
  • Argynnis selene ab. nigromarginata Lambillion, 1906; Rev. Mens. Soc. ent. namur. 6 (10): 48
  • Argynnis selene ab. gerda Schultz, 1908; Ent. Zs. 22 (9): 38
  • Brenthis selene var. castiliana Lowe, 1910; Ent. Rec. J. Var. 22: 23; TL: La Granja
  • Argynnis selene lucetia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 19; TL: Japan
  • interligata (Cabeau, 1919) (Argynnis); Rev. mens. Soc. ent. Namur. 1919: 49
  • halflantsi (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 17
  • consuta (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 50
  • extenuata (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 50
  • Argynnis selene ab. semipallida Nessling, 1925; Notulae Ent. 5 (3): 97; TL: Finland, Vetil, Röyrinki
  • furoshonis (Matsumura, 1929) (Argynnis); Ill. comm. Ins. Japan 1: 16 (repl. sachalinensis Matsumura)
  • Clossiana selene f. h. futura Reuss, 1929; Int. ent. Z. 22: 433
  • medionigrans (Cabeau, 1930) (Argynnis); Lambillionea 30: 54
  • vanescens (Cabeau, 1930) (Argynnis); Lambillionea 30: 54
  • obsoleta (Curtis, 1933) (Argynnis); Trans. ent. Soc. S. Engl. 8: 105
  • Boloria selene (II) gen. postgentilina Verity, 1933; Bull. Soc. ent. Fr. 38 (8): 116
  • Boloria selene r. padimaxima Verity, 1933; Bull. Soc. ent. Fr. 38 (8): 117
  • Brenthis selene ab. striata Lempke, 1936; Tijdschr. Ent. 79: 269; TL: Groningen
  • Argynnis selene ab. e-nigrum Foltin, 1938; Zs. Öst. EntVer. 23: 125; TL: Kefermarkt
  • pseudoselene (Rocci, 1940) (Argynnis); Mem. Soc. ent. ital. 19: 51
  • Argynnis selene ab. novo plesiensis Jesátko, 1944; Acta Soc. ent. bohem. 41: 146
  • Clossiana selene f. loc. hyperborea Lingonblad, 1947; Notulae Ent. 26 (3-4): 69; TL: Finland, Kilpisjärvi
  • Clossiana selene padimaxima f. postpadimaxima Verity, 1950; Le Farfalle diurn. d'Italia 4: 234
  • Argynnis selene ab. fasciata Schnaider, 1950; Polskie Pismo. Ent. 19 (3-4): 241
  • Clossiana selene f. flavescens Lempke, 1956; Tijdschr. Ent. 99 (4): 158
  • Clossiana selene f. debrunneata Lempke, 1956; Tijdschr. Ent. 99 (4): 159
  • Clossiana selene f. infraclara Lempke, 1956; Tijdschr. Ent. 99 (4): 159
  • Clossiana selene f. infradefasciata Lempke, 1956; Tijdschr. Ent. 99 (4): 160
  • Clossiana selene f. semicadmeis Lempke, 1956; Tijdschr. Ent. 99 (4): 160

  • Argynnis selene ab. flavopunctata; Strand, 1912, Ent. Zs. 25 (49): 254
  • Boloria (Clossiana) selene; Pelham, 2008, J. Res. Lepid. 40: 298
  • Boloria selene angustipennis; [LepIndex]; [CoL]
  • Boloria selene castiliana; [LepIndex]; [CoL]
  • Boloria selene debrunneata; [LepIndex]; [CoL]
  • Boloria selene flavescens; [LepIndex]; [CoL]
  • Boloria selene flavopunctata; [LepIndex]; [CoL]
  • Boloria selene furoshonis; [LepIndex]; [CoL]
  • Boloria selene futura; [LepIndex]; [CoL]
  • Boloria selene hyperborea; [LepIndex]; [CoL]
  • Boloria selene infraclara; [LepIndex]; [CoL]
  • Boloria selene infradefasciata; [LepIndex]; [CoL]
  • Boloria selene lucetia; [LepIndex]; [CoL]
  • Boloria selene marphisa; [LepIndex]; [CoL]
  • Boloria selene padimaxima; [LepIndex]; [CoL]
  • Boloria selene postgentilinea; [LepIndex]; [CoL]
  • Boloria selene pseudoselene; [LepIndex]; [CoL]
  • Boloria selene rinaldus; [LepIndex]; [CoL]
  • Boloria selene semicademis; [LepIndex]; [CoL]
  • Boloria selene; [BNA]: 318, #239, pl. 3, 5, 24, 26; [BOC], 281; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [BAMONA] ; [SVF]
  • Brenthis selene var. intermedia; Spångberg, 1878, Bihang K. svenska VetensAkad. Handl. 5: 8
  • Clossiana selene; [SPRK]; [NACL], #4464; [H&R], 104; [BOW]: pl. 2, f. 2-3; [NAB], 559; [EBW]; [BARU], #188; [BRU2]: 45, pl. 32, f. 1-6; Yakovlev, 2012, Nota lepid. 35 (1): 90; [RFEL]; [Mario Maier]

  450x697 (~67Kb) upperside	458x713 (~67Kb) underside	425x374 (~35Kb) upperside
  401x353 (~34Kb) underside	500x391 (~75Kb) upperside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes	500x389 (~57Kb) underside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes
  837x1028 (~120Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio	1371x992 (~199Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio	852x639 (~123Kb) ♀ upperside Russia, Moscow area (36°25'E, 56°00'N), 14.6.2009, Photo © D. Smirnov
  711x624 (~103Kb) ♂ upperside Russia, Moscow area, 14.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov	824x689 (~114Kb) underside Russia, Moscow area, 10.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov	918x694 (~122Kb) ♂ upperside Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
  726x732 (~106Kb) ♂ underside Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov	700x700 (~78Kb) upperside A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin	916x545 (~97Kb) upperside Russia, Moscow area, 15.6.2011 (36°25'E, 56°00'N), Photo © D. Smirnov

  Larva on Viola , V. palustris, V. canina, V. riviniana [SPRK], Fragaria , Vaccinium uliginosum [BRU2]: 46, Viola nephrophylla, V. glabella [BNA]: 318

  • B. s. selene EU, W.Siberia, N.Kazakhstan, Altai, Sayan, Transbaikalia

    • Boloria selene selene; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
    • Clossiana selene selene; [BRU2], 45; [RFEL]
  • B. s. myrina (Cramer, [1777])

    • Papilio myrina Cramer, [1777]; Uitl. Kapellen 2 (9-16): 141, pl. 189, f. B, C
    • Papilio myrinus Herbst, 1798
    • Argynnis myrissa Godart, [1824]; Encyclopédie Méthodique 9 (1): 253
      • = ; Pelham, 2008, J. Res. Lepid. 40: 298
    • Brenthis selene marilandica Clark, 1941; J. Wash. Acad. Sci. 31 (9): 384; TL: Beltsville, Maryland
      • = ; Pelham, 2008, J. Res. Lepid. 40: 298
    • Brenthis myrina ab. nubes Scudder, 1889; Butts eastern U.S. Canada 1: 595; TL: Sanbornton, New Hampshire
      • = ; Pelham, 2008, J. Res. Lepid. 40: 298
    • Brenthis myrina tr.f. nivea Gunder, 1928; Can. Ent. 60 (7): 163 (albino); TL: Staten Island, New York
      • = ; Pelham, 2008, J. Res. Lepid. 40: 299
    • Brenthis selene ab. gunderana Forbes, 1960; Cornell Univ. Agric. Exp. Stn. Mem. 371: 150
      • = ; Pelham, 2008, J. Res. Lepid. 40: 299

    • Argynnis myrina; Scudder, 1863, 166; Scudder, 1868, 379; Edwards, 1876, 161
    • Boloria (Clossiana) myrina myrina; [P&N23]
    • Boloria (Clossiana) myrina; [P&N23]
    • Boloria myrina myrina; [iNaturalist]
    • Boloria myrina; dos Passos & Grey, 1945, 4; [Joseph Belicek]; [MPG] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BOA]
    • Boloria selene marilandica; Klots, 1951, 89; dos Passos, 1964, 89
    • Boloria selene myrina; Klots, 1951, 89; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 245; Pelham, 2008, J. Res. Lepid. 40: 298; [LepIndex]; [CoL]
    • Brenthis myrina; Dyar, 1900, 487; Cary, 1901, 307; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15; Cary, 1906, 439; Elrod, 1906, 69; Franzen, 1914, 366; McDunnough, 1928, 274; Holland, 1931, The Butterfly Book (1931 edn): 105; Clark, 1932, 102; dos Passos & Grey, 1934, 190; McDunnough, 1938, 15; Davenport & Dethier, 1938, 158; Leussler, 1938, 216; Rysgaard, 1939, 193; Comstock, 1940, 53; Brooks, 1942, 34
    • Brenthis myrina ab. nubes; McDunnough, 1938, 16
    • Brenthis selene myrina; Clark, 1941, 384
    • Clossiana myrina nivea; [LepIndex]; [CoL]
    • Clossiana myrina; [LepIndex] ; [CoL]
    • Clossiana selene myrina; [NACL], #4464a; [BNA], #239

    Misidentified/misapplied:

    • myrina (= Papilio bellona); Martyn, 1797, Psyche: pl. 1, f. 2-3 [?]
  • B. s. thalia (Esper, 1789) Arctic Europe, N.Siberia

    • ?Papilio thalia Esper, 1789; Die Schmett., Suppl. Th 1 (3-4): pl. 97, f. 2
    • Papilio thalia Hübner, [1799-1800]; Samml. eur. Schmett. [1]: pl. 11, f. 57-58
    • Argynnis selene var. hela Staudinger, 1861; Stettin Ent. Ztg 22 (10-12): 346
      • = Boloria selene; Pelham, 2008, J. Res. Lepid. 40: 298
    • Boloria selene obscurum Sedykh, 1977; Geogr. Aspect. Conserv. Flora Faun. NE Europ. USSR: 101
      • = Boloria selene; Pelham, 2008, J. Res. Lepid. 40: 298

    • Boloria selene hela; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Boloria selene thalia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [LepIndex]; [CoL]
    • Clossiana selene thalia; [BARU]; [BRU2], 45; [RFEL]
  • B. s. chibiana (Matsumura, 1927) Sakhalin, Kunashir

    • Argynnis selene chibiana Matsumura, 1927; Ins. Matsumurana 2 (2): 116, pl. 3, f. 4; TL: Kuriles

    • Boloria selene chibiana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [LepIndex]; [CoL]
    • Clossiana selene chibiana; [BARU]; [BRU2], 45; [RFEL]
  • B. s. tollandensis (Barnes & Benjamin, 1925) Rocky Mountains (Colorado, Idaho, Montana, New Mexico, Utah, Wyoming), British Columbia

    • Brenthis myrina tollandensis Barnes & Benjamin, 1925; Ent. News 36: 44; TL: Tolland, Gilpin Co., Colorado

    • Boloria (Clossiana) myrina tollandensis; [P&N23]
    • Boloria myrina tollandensis; Brown, 1954, 64; [iNaturalist] ; [BOA]
    • Boloria selene tollandensis; Remington, 1952, 67; Brown, 1957, 64; Albright, 1960, 158; Newcomer & Rogers, 1963, 171; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 253; Pelham, 2008, J. Res. Lepid. 40: 299
    • Brenthis myrina tollandensis; Holland, 1928, 37; Klots, 1937, 328; McDunnough, 1938, 16
    • Brenthis myrina var. tollandensis; Holland, 1931, The Butterfly Book (1931 edn): 106
    • Brenthis selene tollandensis; Clark, 1941, 384
    • Clossiana myrina tollandensis; [LepIndex]; [CoL]
    • Clossiana selene tollandensis; [NACL], #4464d
  • B. s. atrocostalis (Huard, 1927) Nova Scotia, Quebec, Saskatchewan, Manitoba, Maine, Vermont, New Hampshire, New York, Michinag, Wisconsin, Minnesota, North Dakota

    • Argynnis atrocostalis Huard, 1927; Nat. Can. 54 (6): 131; TL: Chicoutini, Quebec
    • Brenthis myrina ab. ♂ serratimarginata Gunder, 1926; Ent. News 37 (1): 7; TL: Vernon, British Columbia
      • = ; Pelham, 2008, J. Res. Lepid. 40: 299
    • Brenthis myrina ab. jenningsae Holland, 1928; Ann. Carnegie Mus. 19 (1): 36; TL: Jellicoe, on Thunder Bay, Ontario
      • = ; Pelham, 2008, J. Res. Lepid. 40: 299
    • Brenthis myrina var. jenningsae Holland, 1931; The Butterfly Book (1931 edn): 105, pl. LVI, f. 9 (type)

    • Boloria (Clossiana) myrina atrocostalis; [P&N23]
    • Boloria myrina atrocostalis; [iNaturalist]
    • Boloria selene atrocostalis; Clark, 1941, 384; Klots, 1951, 89; dos Passos, 1954, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 258, f. 5, 8; [BNA], 318; Pelham, 2008, J. Res. Lepid. 40: 299; [LepIndex]; [CoL]
    • Boloria selene atrocostalis ab. jenningsae; dos Passos, 1964, 89
    • Brenthis myrina atrocostalis; McDunnough, 1930, 107; McDunnough, 1938, 16; dos Passos & Grey, 1934, 190
    • Brenthis myrina atrocostalis ab. jenningsae; McDunnough, 1932, 269; McDunnough, 1938, 16
    • Clossiana selene atrocostalis; [NACL], #4464f
  • B. s. nebraskensis (Holland, 1928) E.Nebraska - Ohio

    • Brenthis myrina var. nebraskensis Holland, 1928; Ann. Carnegie Mus. 19 (1): 36; TL: Dodge Co., Nebraska

    • Boloria (Clossiana) myrina nebraskensis; [P&N23]
    • Boloria myrina nebraskensis; [iNaturalist]
    • Boloria selene nebrascensis; Forbes, 1960, 160 (missp.)
    • Boloria selene nebraskensis; Klots, 1951, 89; Brown, 1957, 64; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 248, f. 2, 6; [BNA], #239; Pelham, 2008, J. Res. Lepid. 40: 299
    • Brenthis myrina nebraskensis; McDunnough, 1938, 16
    • Brenthis myrina r. nebraskensis; Leussler, 1938, 216
    • Brenthis myrina var. nebraskensis; Holland, 1931, The Butterfly Book (1931 edn): 105, pl. LV, f. 8 (type)
    • Brenthis selene nebraskensis; Clark, 1941, 384
    • Clossiana selene nebraskensis; [NACL], #4464b
  • B. s. albequina (Holland, 1928) Yukon, SW.Alaska

    • Brenthis albequina Holland, 1928; Ann. Carnegie Mus. 19 (1): 40, f. 4-6; TL: White Horse Pass, Yukon Territory
    • Brenthis albequina var. baxteri Holland, 1928; Ann. Carnegie Mus. 19 (1): 42; TL: White Horse Pass, Yukon Territory
      • = ; Pelham, 2008, J. Res. Lepid. 40: 299

    • Boloria (Clossiana) myrina albequina; [P&N23]
    • Boloria myrina albequina; [iNaturalist]
    • Boloria selene albequina; dos Passos, 1964, 89; Kohler, 1977, J. Lep. Soc. 31 (4): 255, f. 4; Pelham, 2008, J. Res. Lepid. 40: 299
    • Boloria selene albequina ab. baxteri; dos Passos, 1964, 89
    • Brenthis albequina; Holland, 1931, The Butterfly Book (1931 edn): 107, pl. LV, f. 19-21 (types); McDunnough, 1932, 269
    • Brenthis myrina atrocostalis ab. baxteri; McDunnough, 1938, 16
    • Clossiana albequina baxteri; [LepIndex]; [CoL]
    • Clossiana albequina; [LepIndex] ; [CoL]
    • Clossiana selene albequina; [NACL], #4464e
  • B. s. terraenovae (Holland, 1928) Newfoundland

    • Brenthis myrina terrae-novae Holland, 1928; Ann. Carnegie Mus. 19 (1): 36; TL: Newfoundland

    • Boloria (Clossiana) myrina terraenovae; [P&N23]
    • Boloria myrina terraenovae; [iNaturalist]
    • Boloria selene terrae-novae; Klots, 1951, 89
    • Boloria selene terraenovae; dos Passos, 1964, 89; Shepard, 1975, in Howe, 264; Kohler, 1977, J. Lep. Soc. 31 (4): 262, f. 5; Pelham, 2008, J. Res. Lepid. 40: 299
    • Brenthis myrina terrae-novae; dos Passos, 1935, 87; McDunnough, 1938, 16
    • Brenthis myrina var. terrae-novae; Holland, 1931, The Butterfly Book (1931 edn): 105, pl. LV, f. 13 (m.type)
    • Brenthis selene terrae-novae; Clark, 1941, 384
    • Clossiana myrina terraenovae; [LepIndex]; [CoL]
    • Clossiana selene terraenovae; [NACL]
  • B. s. sugitanii (Seok, 1938) Korea

    • Argynnis selene sugitanii Seok, 1938; Zephyrus 7: 241; TL: Korea

    • Boloria selene sugitanii; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [LepIndex]; [CoL]
    • Clossiana selene sugitanii; [RFEL]
  • B. s. sabulocollis (Kohler, 1977) Nebraska, South Dakota, North Dakota, Colorado

    • Boloria selene sabulocollis Kohler, 1977; J. Lep. Soc. 31 (4): 252, f. 3, 6-7; TL: Smith Lake, Sheridan Co., Nebraska

    • Boloria (Clossiana) myrina sabulocollis; [P&N23]
    • Boloria myrina sabulocollis; [iNaturalist]
    • Boloria selene sabulocollis; Pelham, 2008, J. Res. Lepid. 40: 299; [LepIndex]; [CoL]
    • Clossiana selene sabulicollis[sic, sabulocollis]; [NACL], #4464c
• Boloria perryi (Butler, 1882) S.Ussuri, N.Korea, Amur?. See [maps]

  • Argynnis perryi Butler, 1882; Ann. mag. nat. Hist. (5) 9 (49): 16; TL: Posiette Bay, NE.Corea
  • Argynnis selene var. dilutior Fixsen, 1887; in Romanoff, Mém. Lépid. 3: 303
    • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32

  • Argynnis perrii; [LepIndex] ; [CoL]
  • Boloria perryi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
  • Clossiana perryi; [BRU2]: 46, pl. 32, f. 7-9; [RFEL]
  • Clossiana selene dilutior; [BARU]; [BRU2], 45
  • Clossiana selene perryi; [BARU]
• Boloria jerdoni (Lang, 1868) W.Himalayas, Chitral - Kashmir. See [maps] Jerdon's Silverspot

  • Argynnis jerdoni Lang, 1868; Ent. mon. Mag. 5: 34
  • Argynnis cashmirensis Moore, 1874; Proc. zool. Soc. Lond. 1874 (1): 267, pl. 43, f. 4; TL: Soonamurg, Kashmir
  • Boloria chitralensis Moore, 1900; Lepidoptera Indica 4: 249, pl. 376, f. 3, 3a; TL: Kashmir

  • Argynnis jerdoni; [BIR], 223
  • Clossiana jerdoni chitralensis; [LepIndex]; [CoL]
  • Clossiana jerdoni; [BOW]: pl. 194, f. 13; [LepIndex] ; [CoL]
• Boloria erubescens (Staudinger, 1901) -. See [maps]

  • Argynnis hegemone var. erubescens Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 35; TL: Korla

  • Boloria erubescens; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
  • Clossiana erubescens; [Oleg Kosterin]; [LepIndex] ; [CoL] ; [LEPIFORUM]
  • B. e. erubescens

    • Boloria erubescens erubescens; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
  • B. e. tienschanica (Wagner, 1913) Tian-Shan

    • Argynynnis hegemone tienschanica Wagner, 1913; Ent. Mitt. 2 (4): 114, f. 9; TL: Juldus
      • = Clossiana hegemone hegemone; [BRU2], 46

    • Clossiana erubescens tienschanica; [Oleg Kosterin]

    400x533 (~49Kb) underside An alpine meadow on the right bank terrace of the fourth tributary of the Sargir River (the Usek River basin), altitude 3200 m, 47 km NEN of Zharkent, 21 km NEN of village Sarybel', the mountain massif Tyshkantau, the Southern Dzhungarian Alatau mountain chain (NE part of Tien Shan), Zharkent District, Taldy-Kurgan Province, SE Kazakhstan. 27th of July 1994, Photo © Oleg Kosterin

  • B. e. chotana (Bang-Haas, 1915) Pakistan, ...

    • Argynnis hegemone var. chotana Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3): 100; TL: Kun-Lun Mts., China
    • Clossiana franciscana Verhulst, 1999; Lambillionea 99 (2): 213; TL: Pakistan, Gilgit
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossia hegemone ali Churkin, 2002; Helios 3: 162; TL: Alai
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32

    • Boloria erubescens chotana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana erubescens chotana; [LepIndex]; [CoL]
    • ?Clossiana hegemone chotana; [BRU2], 46 (?)
  • B. e. haberhaueri Hemming, 1933 Alatau, Margelan

    • Argynnis hegemone Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 292 (preocc. Argynnis hegemone Godart, 1819); TL: "Ala Tau" [Dzhungarsky Alatau Mts., Kazakhstan]
    • Argynnis hegemone Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 407, pl. 15, f. 16-17 (Staudinger i.l.); TL: Kuldja?
    • Boloria haberhaueri Hemming, 1933; Entomologist 66 (12): 275 (repl. Argynnis hegemone Staudinger, 1881)

    • Argynnis hegemone; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 437
    • Clossiana erubescens haberhaueri; [LepIndex]; [CoL]
    • Clossiana hegemone; [BOW]: pl. 194, f. 12; [BRU2]: 46, pl. 33, f. 25-30
  • B. e. houri Wyatt, 1961 W.Tian-Shan, Ghissar-Darvaz, Pamirs-Alai

    • Boloria hegemone houri Wyatt, 1961; J. Lep. Soc. 15 (1): 8; TL: Bala Quran, Anjuman Valley, Hindu Kush Mts., 14300ft

    • Boloria erubescens houri; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana erubescens houri; [LepIndex]; [CoL]
    • Clossiana hegemone houri; [BRU2], 46
  • B. e. eximia (Churkin, 2002) Fergansky Mts

    • Clossiana hegemone eximia Churkin, 2002; Helios 3: 164; TL: Fergansky Mts, 15km E Kalmak-Krchin v., Kugart R., 2000-2200m

    • Boloria erubescens eximia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
  • B. e. tuzovi (Churkin, 2002) Talassky Alatau

    • Clossiana hegemone tuzovi Churkin, 2002; Helios 3: 165; TL: Talassky Alatau, Alabel Pass, 3300m

    • Boloria erubescens tuzovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
  • B. e. ancilla (Churkin, 2004) Tadjikistan

    • Clossiana hegemone ancilla Churkin, 2004; Helios 5: 132; TL: Tadjikistan, Peter the Great Range, 23km SE Tadzhikobad, Ganishou v.
      • = Boloria erubescens houri; Korb & Bolshakov, 2016, Zootaxa 4160 (1): (1-234)

    • Boloria erubescens ancilla; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
  • B. e. panterai Korb, 2005

    • Boloria erubescens panterai Korb, 2005; A catalogue of butterflies of the ex-USSR: 86
• Boloria selenis (Eversmann, 1837) -. See [maps]

  • Argynnis selenis Eversmann, 1837; Bull. Soc. imp. Nat. Moscou 1837 (1): 10; TL: "Kasan" [Kazan, Tatarstan, Russia]
  • ?Clossiana speranda Grosser, 1979; Reichenbachia 17 (39): 331; TL: NW.Mongolia
    • = ?Clossiana selenis; [BRU2], 44 (?)
  • Argynnis selenis takamukuella Matsumura, 1929; Insecta Matsumurana 3 (4): 155; TL: Korea
  • mariapaula (Fernández Vidal, 1980) (Clossiana); Revta lepid. 8 (3): 215

  • Boloria selenis; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist] ; [LEPIFORUM]
  • Clossiana selenis mariapaula; [LepIndex]; [CoL]
  • Clossiana selenis takamukuella; [LepIndex]; [CoL]
  • Clossiana selenis; [BARU], #189; [BOW]: pl. 194, f. 15; [MRS], 478; [BRU2]: 43, pl. 31, f. 25-30; [RFEL]; [LepIndex] ; [CoL]
  • Clossiana speranda; [LepIndex] ; [CoL]

  Larva on Viola [BRU2]: 44

  • B. s. selenis S.Urals, Volga Valley

    • Boloria selenis selenis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
    • Clossiana selenis selenis; [BRU2], 43
  • B. s. sibirica (Erschoff, 1870) Amur, NE.Siberia

    • Argynnis selenis var. sibirica Erschoff, 1870; Bull. Soc. Imp. Nat. Moscou 43 (1): 112
    • Argynnis selenis var. festiva Krulikovsky, 1893; Soc. Ent. 7 (21) (23): 182; TL: Siberia orientalis
    • Argynnis selenis chosensis Matsumura, 1927; Insecta Matsumurana 1 (4): 165, pl. 5, f. 6; TL: Corea, Shuotsu
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Argynnis (Brenthis) selenis meinhardi Sheljuzhko, 1929; Mitt. Münch. Ent. Ges. 19: 359; TL: Tomsk
    • Brenthis selenis kononovi Kurentzov, 1970; Butterflies of the USSR Far East: 102
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32

    • Boloria selene meinhardi; [LepIndex]; [CoL]
    • Boloria selenis sibirica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
    • Clossiana selenis chosensis; [BRU2], 44; [LepIndex]; [CoL]
    • Clossiana selenis kononovi; [BRU2], 44
    • Clossiana selenis sibirica; [Oleg Kosterin]; [BRU2], 44; Yakovlev, 2012, Nota lepid. 35 (1): 90; [RFEL]; [LepIndex]; [CoL]

    532x326 (~51Kb) underside A southern steppefied slope of the mountain on the Chuya River right bank between the villages Kurai and Chagan-Uzun, Altai Mts., West SIberia, Russia. 7th July 1988, Photo © Oleg Kosterin

  • B. s. onorensis (Matsumura, 1925) Sakhalin

    • Argynnis selenis onorensis Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ. 15: 100, pl. 8, f. 1 ♂; TL: Saghalin

    • Boloria selene onorensis; [LepIndex]; [CoL]
    • Clossiana selenis onorensis; [BRU2], 44
• Boloria oscarus (Eversmann, 1844) -. See [maps]

  • Argynnis oscarus Eversmann, 1844; Bull. Soc. Imp. Nat. Moscou 17 (3): 588, pl. 14, f. 1 a,b; TL: Altai [Irkutsk region?]
  • mandarina (Matsumura, 1939) (Argynnis); Bull. biogeogr. Soc. Japan 9 (20): 355; TL: Manchuria

  • Boloria oscarus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist] ; [LEPIFORUM]
  • Clossiana oscarus mandarina; [LepIndex]; [CoL]
  • Clossiana oscarus; [BARU], #192; [BOW]: pl. 194, f. 14; [BRU2]: 44, pl. 34, f. 1-6; [RFEL]; [LepIndex] ; [CoL]

  488x400 (~38Kb) ♂ underside A bushy meadow on the second right bank terrace of the Koen River at the Volchikha Rivulet just upstream of its fall into the Koen River near the village Nizhniy Koen, Iskitim District, Novosibirsk Province, West Siberia, Russia. 18th June, 1989, Photo © Oleg Kosterin

  Larva on Viola [BARU]

  • B. o. oscarus S.Siberia, E.Amur

    • Boloria oscarus oscarus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist]
    • Clossiana oscarus oscarus; [BRU2], 44; [RFEL]
  • B. o. australis (Graeser, 1888) S.Amur, Ussuri, Sakhalin

    • Argynnis oscarus var. australis Graeser, 1888; Berl. Ent. Zs. 32 (1): 90; TL: "bei Wlad."
    • Argynnis oscarus f.[?] major Graeser, 1888; Berl. Ent. Zs. 32 (1): 90
    • Argynnis clossiana var. maxima Fixsen, 1887; in Romanoff, Mém. Lépid. 3: 304 (preocc.)
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33

    • Boloria oscarus australis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [RFEL]
    • Clossiana oscarus australis; [BARU]; [BRU2], 44; [LepIndex]; [CoL]
  • B. o. oscaroides (Ménétriés, 1859) E.Yakutia

    • Argynnis oscarus (?) var. oscaroides Ménétriés, 1859; Bull. phys.-mat. Acad. Sci. St. Pétersb. 17 (31): 497, Mélanges biol. St.-Pétersb. 3 (2):216, pl. 5, f. 3-4

    • Boloria oscarus oscaroides; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana oscarus oscaroides; [BRU2], 44; [RFEL]
• Boloria euphrosyne (Linnaeus, 1758) . See [maps] Perlada rojiza Skogspärlefjäril Silberstrich Dostojka eufrozyna Pursuhopeatälä Pearl-bordered Fritillary Prydlig pärlemorfjäril Grand Collier Argenté

  • Papilio euphrosyne Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 481, no. 142
  • Papilio euphrosyne Linnaeus, 1761; Fauna Suecica (Edn 2): 282; TL: Sweden
  • Papilio argenticollis Retzius, 1783; Gen. Spec. Ins.: 31
  • Brenthis morrisii Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3): 246; TL: California [error]
    • = Argynnis euphrosyne; Strecker, 1878, Synon. Cat. Macroplep.: 118
    • = Clossiana euprosyne; Pelham, 2008, J. Res. Lepid. 40: 442
  • Brenthis euphrosyne ab. obsoleta Tutt, 1896; Brit. Butts: 300
  • Brenthis euphrosyne ab. pauca Tutt, 1896; Brit. Butts: 300
  • Argynnis euphrosyne v. apennina Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 35
  • Argynnis euphrosyne ab. melanotica Spuler, 1901; Schmett. Eur. 1: 27
  • Argynnis euphrosyne ab. pallida Spuler, 1901; Schmett. Eur. 1: 27
  • Argynnis euphrosyne ab. radiata Spuler, 1901; Schmett. Eur. 1: 27
  • Argynnis euphrosyne ab. tatrica Aigner-Abafi, 1906; Ann. hist.-nat. Mus. Hung. 4 (2): 502; TL: Tatra
  • Argynnis euphrosyne f. answina Fruhstorfer, 1907; Ent. Zs. 21 (27): 164
  • Argynnis euphrosyne fingal f. niveola Stichel, 1908; Berl. ent. Z. 53 (1): 81, pl. 3, f. 4
  • Argynnis euphrosyne obscurior Seitz, 1909; Gross-Schmett. Erde 1: 229, pl. 67 h (Stgr. i.l.)
  • Argynnis euphrosyne var. phaenna Jachontov, 1909; Revue Russe Ent. 8: 285; TL: Transcaucasia
  • Argynnis euphrosyne cynosoma Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 15; TL: Salève
  • Argynnis euphrosyne calynde Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 16; TL: Tessin, Fusio, Val Piora, 1250m
  • Argynnis euphrosyne neston Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 16; TL: Tessin, Mt Generoso
  • interligata (Cabeau, 1919) (Argynnis); Rev. mens. Soc. ent. Namur. 1919: 49
  • chlorographa (Cabeau, 1920) (Argynnis); Rev. mens. Soc. ent. Namur. 1920: 21; TL: France
  • capronnieri (Derenne, 1922) (Argynnis); Rev. Mens. Soc. ent. Namur. 1922: 36
  • rabesina (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 1
  • Brenthis euphrosyne ab. plumbea Cockayne, 1924; Ent. Rec. J. Var. 36: 8; TL: Chilterns
  • Brenthis euphrosyne alpina Ebert, 1926; Dt. ent. Z. Iris 40: 33
  • Argynnis euphrosyne ab. pittionii Nitsche, 1926; Verh. zool.-bot. Ges. 74/75 (s.b.): 17
  • forstorpensis (Strand, 1927) (Argynnis); Ent. Tidskr. 48: 56
  • Boloria euphrosyne r. nestonclara Verity, 1932; Ent. Rec. J. Var. 44: 112; TL: Germany
  • Boloria euphrosyne r. padimira Verity, 1932; Ent. Rec. J. Var. 44: 113; TL: Apennines
  • Boloria euphrosyne r. eminens Verity, 1932; Ent. Rec. J. Var. 44: 113; TL: Ural Mts
  • Boloria euphrosyne r. variana Verity, 1932; Ent. Rec. J. Var. 44: 114; TL: England
  • Boloria euphrosyne r. magnaclara Verity, 1932; Ent. Rec. J. Var. 44: 120; TL: Amasia
  • Boloria euphrosyne r. orientisvivax Verity, 1932; Ent. Rec. J. Var. 44: 121; TL: Sajan Mts
  • vittata (Nordström, 1933) (Brenthis); Ent. Tidskr. 54: 175; TL: Lapland
  • lecharlesi (Varin, 1933) (Argynnis); Amat. Papill. 6: 233
  • septentrionalis (Nordström, 1933) (Brenthis); Ent. Tidskr. 54: 171; TL: Lapland
  • Argynnis euphrosyne lavinia Hemming, 1934; Stylops 3 (9): 193 (repl. Brenthis euphrosyne alpina Ebert, 1926)
  • dorensis (Réal, 1950) (Argynnis); Rev. franc. Lép. 12: 214; TL: France
  • Clossiana euphrosyne r. austreminens Verity, 1950; Le Farfalle diurn. d'Italia 4: 240
  • Clossiana euphrosyne neston sr. austroneston Verity, 1950; Le Farfalle diurn. d'Italia 4: 240
  • Argynnis euphrosyne ab. edna Lobb, 1953; Ent. Rec. J. Var. 65: 56, pl. 3, f. 4-5; TL: Parkhurst Forest, Isle of Wight
  • Brenthis euphrosyne esperi Heydemann, 1955; Scr. Naturw. Ver. Schleswig-Holstein 27 (2): 136 (Esper, 1777 1 (1) pl. 18)
  • galiseri (Niculescu, 1957) (Argynnis); Comun. Acad. Repub. Rom. 7: 456
  • albomaculata (Goodson, 1959) (Clossiana); Entomologist 92: 148

  • Boloria (Clossiana) euphrosyne; [P&N23]
  • Boloria euphrosyne; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SVF]
  • Clossiana euphrosyne answina; [LepIndex]; [CoL]
  • Clossiana euphrosyne apennina; [LepIndex]; [CoL]
  • Clossiana euphrosyne austreminens; [LepIndex]; [CoL]
  • Clossiana euphrosyne calynde; [LepIndex]; [CoL]
  • Clossiana euphrosyne cynosoma; [LepIndex]; [CoL]
  • Clossiana euphrosyne eminens; [LepIndex]; [CoL]
  • Clossiana euphrosyne lavinia; [LepIndex]; [CoL]
  • Clossiana euphrosyne magnaclara; [LepIndex]; [CoL]
  • Clossiana euphrosyne neston; [LepIndex]; [CoL]
  • Clossiana euphrosyne nestonclara; [LepIndex]; [CoL]
  • Clossiana euphrosyne niveola; [LepIndex]; [CoL]
  • Clossiana euphrosyne obscurior; [LepIndex]; [CoL]
  • Clossiana euphrosyne orientisvivax; [LepIndex]; [CoL]
  • Clossiana euphrosyne padimira; [LepIndex]; [CoL]
  • Clossiana euphrosyne phaenna; [LepIndex]; [CoL]
  • Clossiana euphrosyne pittionii; [LepIndex]; [CoL]
  • Clossiana euphrosyne septentrionalis; [LepIndex]; [CoL]
  • Clossiana euphrosyne; [SPRK]; [H&R], 105; [BOW]: pl. 1, f. 26; [EBW]; [BARU], #190; [BRU2]: 44, pl. 31, f. 13-24; [RFEL]; [LepIndex] ; [CoL] ; [Mario Maier]

  500x349 (~71Kb) upperside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes	417x500 (~57Kb) underside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes	533x400 (~50Kb) larva An open pine grove above the bluff of the valley of the Bol'shoi Elbash River, 5 km NW of the village Ust'-Chem, Iskitim dist., Novosibirsk region, West Siberia, 9th May 1995 (mature larva), Photo © Oleg Kosterin
  600x832 (~62Kb) underside Finland, Loppi, Luutasuo 673:35, 2.6.1979, Photo © Pertti Pakkanen	900x745 (~146Kb) ♀ upperside FINLAND: Li: Inari, Luttojoki, 760:54, 11.7.1998, Photo © Markku Savela	600x722 (~61Kb) upperside FINLAND, Ab: Suomusjärvi, 669:31, 15.6.1997, Photo © Markku Savela
  900x1291 (~125Kb) Finland, Ab: Suomusjärvi (669:31), 6.6.2004, Photo © Markku Savela	1258x972 (~356Kb) ♂ upperside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Republic, West Siberia, Russia. 30th June 2007, Photo © Oleg Kosterin	861x1128 (~219Kb) ♂ underside An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 2nd July 2007, Photo © Oleg Kosterin
  779x555 (~89Kb) upperside Russia, Moscow area, 11.6.2011 (38°52'E, 55°33'N), Photo © D. Smirnov	632x611 (~109Kb) underside Russia, Moscow area, 11.6.2011 (38°52'E, 55°33'N), Photo © D. Smirnov	800x631 (~57Kb) underside FINLAND: Ka: Virolahti, 671:53 (Reskopinsuo), 14.6.1997, Photo © Markku Savela

  Larva on Viola , V. palustris, V. riviniana, Vaccinium uliginosum, Ledum palustre [SPRK], Viola hirta, Waldsteinia ternata [BRU2]: 45

  • B. e. euphrosyne CEU, Siberia

    • Argynnis euphrosyne euphrosyne; Fruhstorfer, 1907, Ent. Zs. 21 (27): 164
    • Boloria euphrosyne euphrosyne; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
    • Clossiana euphrosyne euphrosyne; [BRU2], 45; [RFEL]
  • B. e. fingal (Herbst, 1800) NEU, Siberia

    • euphrosyne fingal (Herbst, 1800) (Papilio); Naturs. Schmett. 10: pl. 270, f. 1-3
      • = Boloria euphrosyne lapponica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Papilio lapponica Esper, 1798; Die Schmett., Suppl. Th 1 ([7]): 83, pl. 108, f. 5

    • Argynnis euphrosyne fingal; Fruhstorfer, 1907, Ent. Zs. 21 (27): 164
    • Boloria euphrosyne lapponica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana euphrosyne fingal; [BRU2], 45; [LepIndex]; [CoL]
  • ?B. e. nephele (Herrich-Schäffer, [1847]) Ural, Siberia

    • euphrosyne nephele (Herrich-Schäffer, [1847]) (Argynnis); Syst. Bearb. Schmett. Europ. 1 (26): (ii) f. 371-372

    • Argynnis euphrosyne nephele; Fruhstorfer, 1907, Ent. Zs. 21 (27): 164
    • Clossiana euphrosyne nephele; [BARU]
  • B. e. dagestanica (Sovinsky, 1905) Transcaucasia

    • Argynnis euphrosyne var. dagestanica Sovinsky, 1905; Revue Russe Ent. 5 (3-4): 108

    • Boloria euphrosyne dagestanica; [iNaturalist]
    • Clossiana euphrosyne dagestanica; [LepIndex]; [CoL]
  • B. e. anka (Fruhstorfer, 1907) Caucasus

    • Argynnis euphrosyne anka Fruhstorfer, 1907; Ent. Zs. 21 (27): 164; TL: Caucasus
    • Clossiana phaenea Jachontov, 1908

    • Boloria euphrosyne anka; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana euphrosyne anka; [LepIndex]; [CoL]
  • B. e. orphana (Fruhstorfer, 1907) Transbaikalia, Amur, Ussuri

    • Argynnis euphrosyne orphanus Fruhstorfer, 1907; Ent. Zs. 21 (27): 164

    • Clossiana euphrosyne orphana; [BRU2], 45
    • Clossiana euphrosyne orphanus; [BARU]; [LepIndex]; [CoL]
  • B. e. rusalka (Fruhstorfer, 1909) SEU, W.Siberia

    • Argynnis euphrosyne rusalka Fruhstorfer, 1909; Int. ent. Zs. 3 (20): 112; TL: Saratow

    • Boloria euphrosyne rusalka; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana euphrosyne rusalka; [BRU2], 45; [LepIndex]; [CoL]
  • B. e. densoi (Fruhstorfer, 1909)

    • Argynnis euphrosyne densoi Fruhstorfer, 1909; Int. ent. Zs. 3 (20): 113; TL: Champery

    • Clossiana euphrosyne densoi; [LepIndex]; [CoL]
  • B. e. kamtschadalis (Seitz, 1909) Kamchatka, N.Sakhalin

    • Argynnis euphrosyne kamtschadalis Seitz, 1909; Gross-Schmett. Erde 1: 230, pl. 67 h (Stgr. i.l.); TL: Kamchatka

    • Boloria euphrosyne kamtschadalus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana euphrosyne kamtschadalis; [LepIndex]; [CoL]
    • Clossiana euphrosyne kamtschadalus; [BARU]; [BRU2], 45; [RFEL]
  • B. e. umbra (Seitz, 1909) Altai, Sayan

    • Argynnis euphrosyne f. umbra Seitz, 1909; Gross-Schmett. Erde 1: 229, pl. 67 h

    • Clossiana euphrosyne umbra; [BRU2], 45; Yakovlev, 2012, Nota lepid. 35 (1): 90; [LepIndex]; [CoL]
  • B. e. orphanoides (Huang & Murayama, 1992) Xinjiang

    • Clossiana euphrosyne orphanoides Huang & Murayama, 1992; Tyô to Ga 43 (1): 7, f. 17; TL: Jiadengyu, Altai, 1380-1600m

    • Boloria euphrosyne orphanoides; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
• Boloria iphigenia (Graeser, 1888) -. See [maps]

  • Argynnis iphigenia Graeser, 1888; Berl. Ent. Zs. 32 (1): 90; TL: Amur "downstream of Khabarovsk"

  • Boloria iphigenia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32; [iNaturalist]
  • Clossiana iphigenia; [BARU], #191; [BRU2]: 45, pl. 32, f. 10-12; [RFEL]; [LepIndex] ; [CoL]

  Larva on Viola selkirkii, (mainland,Japan) Viola grypoceras, (Sakhalin,Japan) [BRU2]: 45

  • B. i. iphigenia E.Amur, Ussuri, NE.China

    • Boloria iphigenia iphigenia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana iphigenia iphigenia; [BRU2], 45; [RFEL]
  • B. i. sachalinensis (Matsumura, 1908) Japan

    • Argynnis sachalinensis Matsumura, 1908; Ent. Zs. 22 (39): 158:Hokkaido

    • Argynnis euphrosyne sachalinensis; Esaki, 1916, Ent. Mag. Kyoto 2 (1): 48, 50, pl. 2, f. 6; Matsumura, 1925, J. Coll. Agric. Hokkaido imp. Univ. 15: 99
    • Argynnis euphrosyne var. sachalinensis; Matsumura, 1919, Thous. Ins. Japan. Addit. 3: 585, pl. 45, f. 10
    • Argynnis selene sachalinensis; Nire, 1916, Zool. Mag. Tokyo 28: 64
    • Boloria selene sachalinensis; [LepIndex]
    • Clossiana euphrosyne sachalinensis; [LepIndex]; [CoL]
    • Clossiana iphigenia sachalinensis; Kudrna, 1974, Atalanta 5: 108; [LepIndex]; [CoL]
    • ?Clossiana iphigenia sachalinensis; [BARU]
  • B. i. insularia (Korshunov, 1996) Sakhalin, Kunashir

    • iphigenia insularia (Korshunov, 1996) (Clossiana); Dopolneniya i ispravleniya k knige "Dnevnye babochki aziatskoi chasti Rossii": [?] (repl. Argynnis sachalinensis Matsumura, 1936 ?? ); TL: Sakhalin
    • Clossiana iphigenia alpharatoria Korb, 1997
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32

    • Boloria iphigenia insularia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
    • Clossiana iphigenia alpharatoria; [BRU2], 45
    • Clossiana iphigenia insularia; [RFEL]
• Boloria kriemhild (Strecker, 1879) Montana, Wyoming. See [maps] Kriemhild Fritillary Relict Fritillary

  • Argynnis kriemhild Strecker, 1879; in McCauley, Rept. San Juan Recon. 5 (1) pt. 2 (3) appendix SS: 1854, pl. 1, f. 5-6
  • Argynnis laurenti Skinner, 1913; Ent. News 24 (10): 450; TL: Silver Lake, Utah
    • = ; Pelham, 2008, J. Res. Lepid. 40: 301
  • Clossiana lücki [= luecki] Reuss, 1923; Soc. Ent. 38 (4): 14; TL: Utah
    • = ; Pelham, 2008, J. Res. Lepid. 40: 301

  • Argynnis laurenti; [LepIndex] ; [CoL]
  • Boloria (Clossiana) kriemhild; Pelham, 2008, J. Res. Lepid. 40: 301; [P&N23]
  • Boloria epithore kriemhild; [LepIndex]; [CoL]
  • Boloria kriemhild; [BNA]: 320, #241, pl. 26; [Opler]; [MPG] ; [iNaturalist] ; [BOA] ; [BAMONA]
  • Brenthis epithore kreimhild[sic]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • Clossiana euphrosyne kriemhild; [LepIndex]; [CoL]
  • Clossiana kriemeld; McHenry, 1964, (missp.)
  • Clossiana kriemhild; [NACL], #4468; [NAB], 564
  • Clossiana lucki; dos Passos, 1964, (missp.); [LepIndex] ; [CoL]

  Larva on Viola [NACL]

• Boloria freija (Thunberg, 1791) . See [maps] Myrpärlefjäril Zigzag Fritillary Freya's Fritillary Freija's Fritillary Frejas pärlemorfjäril Muurainhopeatäplä

  • Papilio freija Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 34; TL: Vestrobothnia & Lapponica
  • ?Papilio freija Beclin, 1791
  • Argynnis freya[sic] ab. obscura Schöyen, 1885; Ent. Tidskr. 6: 142
  • Brenthis freija ab. fasciata Sheldon, 1912; Entomologist 45 (585): 66
  • ?daisetuzana Matsumura

  • Boloria (Clossiana) freija; Pelham, 2008, J. Res. Lepid. 40: 304
  • Boloria freija; [BNA]: 323, #249, pl. 26; [BOC], 284; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [MPG] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [BOA] ; [BAMONA] ; [SVF]
  • Brenthis freija; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • Clossiana freija daisetuzana; [LepIndex]; [CoL]
  • Clossiana freija; [SPRK]; [H&R], 106; [BARU], #201; [NACL], #4471; [MRS], 477; [NAB], 566; [BRU2]: 48, pl. 32, f. 24-29; [RFEL]; [LepIndex] ; [CoL]

  Larva on Rubus chamaemorus, Vaccinium uliginosum [SPRK], Vaccinium vitis-idaeus, Rhododendron auerum [BARU], Empetrum nigrum [MRS], Vaccinium caespitosum, V. uliginosum, Arctostaphylos uva-ursi, Empetrum nigrum [NAB], Arctous alpina, (Scandinavia) Sieversia , Rhododendron , (Japan) Dryas integrifolia, (N.America) [BRU2]: 48

  • B. f. freija Fennoscandia, W.Siberia, C.Siberia, Far East, Transbaikiali, Chukotka, Amur, Ussuri, Canada, Alaska

    • Argynnis freya Godart, 1819; Encyclopédie Méthodique 9 (1): 273 (repl. Argynnis freija Thunberg, 1791)
      • = ; Pelham, 2008, J. Res. Lepid. 40: 304
    • Papilio freija Quensel, 1791; K. VetenskAcad. Handl. [2] 12: 276, pl. 10, f. 5-6 (preocc. Papilio freija Thunberg, 1791)
    • Papilio lapponica Esper, 1789; Die Schmett., Suppl. Th 1 (3-4): 21, pl. 97, f. 3
      • = ; Pelham, 2008, J. Res. Lepid. 40: 304
    • Argynnis (Bretis[sic]) freija jakutensis Wnukowsky, 1927; Mitt. Münch. Ent. Ges. 17 (7-12): 70; TL: Jakutia
      • = ; Pelham, 2008, J. Res. Lepid. 40: 304
      • = Clossiana freija; [RFEL]
    • Brenthis freija tr.f. gunderi Harper, 1933; Pan-Pacific Ent. 9 (3): 99; TL: McCreary, Manitoba, Canada
      • = ; Pelham, 2008, J. Res. Lepid. 40: 304
    • Clossiana freija zamolodchikova Churkin, 2001; Helios 2: 99, pl. 8, f. 10-12
      • = ; Pelham, 2008, J. Res. Lepid. 40: 304
      • = Boloria freija jakutensis (zamolodchikovi); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
      • = Clossiana freija (zamolodchikovi); [RFEL]

    • Argynnis lapponica; [LepIndex] ; [CoL]
    • Boloria (Clossiana) freija freija; [P&N23]
    • Boloria (Clossiana) freija; [P&N23]
    • Boloria freija freija; [BNA], #249; Pelham, 2008, J. Res. Lepid. 40: 304; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist] ; [BOA]
    • Boloria freija jakutensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist]
    • Clossiana freija freija; [BRU2], 48; [CoL]
    • Clossiana freija gunderi; [LepIndex]; [CoL]
    • Clossiana freija jakutensis; [BARU]; [BRU2], 48; [LepIndex]; [CoL]
  • B. f. tarquinius (Curtis, 1835) British Columbia, ...

    • Melitaea tarquinius Curtis, 1835; in Ross, Nar. 2nd Voy. N.W. Passage: lxviii

    • Argynnis lapponica tarquinius; [LepIndex]; [CoL]
    • Boloria (Clossiana) freija tarquinius; [P&N23]
    • Boloria freija tarquinius; Pelham, 2008, J. Res. Lepid. 40: 304; [iNaturalist] ; [BOA]
    • Brenthis freija tarquinius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
    • Clossiana freija tarquinius; [NACL], #4471a; [LepIndex]; [CoL]
  • B. f. pallida (Elwes, 1899) Altai, Sayan, Tuva

    • Argynnis freija var. pallida Elwes, 1899; Trans. ent. Soc. Lond. 1899 (3): 338 (?preocc. Argynnis aglaja pallida Tutt, 1896)
    • Boloria freija calais Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 17; TL: Schawyr, Tannuola

    • Boloria freija calais; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana freija calais; [BARU](Tuva) , ; [LepIndex]; [CoL]
    • Clossiana freija pallida; [BARU]; [BRU2], 48; [Oleg Kosterin]; Yakovlev, 2012, Nota lepid. 35 (1): 90; [LepIndex]; [CoL]

    533x400 (~67Kb) ♀ underside A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 13th July, 1988, Photo © Oleg Kosterin

    508x444 (~38Kb) ♂ upperside A meadowy-tundrous SW slope of a ridge between the headwaters of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Aster alpinus), Photo © Oleg Kosterin

    600x600 (~71Kb) ♂ upperside A meadowy-tundrous SW slope of a ridge between the headwaters of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Aster alpinus), Photo © Oleg Kosterin

  • B. f. asahidakeana (Matsumura, 1926) Japan

    • Argynnis asahidakeana Matsumura, 1926; Insecta Matsumurana 1 (2): 104; TL: Hokkaido, Mt Daisetsu

    • Argynnis asahidakeana; [LepIndex] ; [CoL]
    • Boloria freija asahidakeana; [iNaturalist]
    • Clossiana freija asahidakeana; Kudrna, 1974, Atalanta 5: 109
  • B. f. browni (Higgins, 1953) Rocky Mountains

    • Clossiana freija browni Higgins, 1953; Entomologist 86 (9): 210

    • Boloria (Clossiana) freija browni; [P&N23]
    • Boloria freija browni; [BNA], #249; Pelham, 2008, J. Res. Lepid. 40: 304; [iNaturalist] ; [BOA]
    • Clossiana freija browni; [NACL], #4471d; [LepIndex]; [CoL]
• Boloria natazhati (Gibson, 1920) Yukon, Victoria I.. See [maps] Pleistocene Fritillary

  • Brenthis natazhati Gibson, 1920; Rep. Can. arctic exped. 1913-18. 3 (1): 21 i; TL: Mt. Natazhat, St. Elias Mts., Yukon Territory, 2470-2620m
  • Boloria natazhati perunovi Korb, 1999; Lambillionea 99 (4): (569-571)

  • Boloria (Clossiana) natazhati natazhati; [P&N23]
  • Boloria (Clossiana) natazhati; Pelham, 2008, J. Res. Lepid. 40: 304; [P&N23]
  • Boloria freija natazhati; dos Passos, 1964; Howe, 1975
  • Boloria natazhati; [BNA]: 323, #250, pl. 24; Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 181, f. 1-2; [Opler]; [MPG] ; [iNaturalist] ; [BOA] ; [BAMONA]
  • Clossiana freija natazhati; Miller & Brown, 1981; Ferris, dos Passos, Ebner & Lafontaine, 1983; [NACL], #4471b; Tilden & Smith, 1968
  • Clossiana natazhati; [LepIndex] ; [CoL]

  Larva on Dryas integrifolia? Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 181

  • B. n. natazhati

    • Boloria natazhati natazhati; Pelham, 2008, J. Res. Lepid. 40: 304; [iNaturalist] ; [BOA]
  • B. n. nabokovi Stallings & Turner, 1947 Alaska

    • Boloria freija nabokovi Stallings & Turner, 1947; Can. Ent. 78 (7-8): 134; TL: Alaska Military Highway, 102 miles N of Summit 2, 1830m
      • = Boloria natazhati; Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 181

    • Boloria (Clossiana) natazhati nabokovi; [P&N23]
    • Boloria freija nabokovi; dos Passos, 1964; Howe, 1975
    • Boloria natazhati nabokovi; Pelham, 2008, J. Res. Lepid. 40: 305; [iNaturalist] ; [BOA]
    • Clossiana freija nabokovi; Miller & Brown, 1981; [NACL], #4471c; Tilden & Smith, 1986; [LepIndex]; [CoL]
  • B. n. bankslandia Shepard, 1998 Victoria Is.

    • Boloria natazhati bankslandia Shepard, 1998; in Emmel, Systematics W. North Am. Butts.: 732

    • Boloria (Clossiana) natazhati bankslandia; [P&N23]
    • Boloria natazhati bankslandia; Pelham, 2008, J. Res. Lepid. 40: 305; [iNaturalist] ; [BOA]
• Boloria angarensis (Erschoff, 1870) -. See [maps]

  • Argynnis angarensis Erschoff, 1870; Bull. Soc. Imp. Nat. Moscou 43 (1): 112; TL: "the Irkutsk Province"

  • Boloria angarensis; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist] ; [LEPIFORUM]
  • Clossiana angarensis; [BARU], #193; [BOW]: pl. 194, f. 10; [BRU2]: 43, pl. 30, f. 7-19; [RFEL]; [LepIndex] ; [CoL]

  533x400 (~45Kb) ♀ upperside An open larch (Larix gmelinii (Rupr.) Rupr.)/birch (Betula pendula Roth. and B. davurica Pallas) forest above the left bluff of the Budyumkan River valley 5 km of its confluence with the Argun' River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 2nd August 1997, Photo © Oleg Kosterin

  352x299 (~22Kb) ♀ underside A road in a larch (Larix gmelinii (Rupr.) Rupr.) forest not far from the Argun' River left bank between the village Uryupino and the Gazimur River, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 31th July 1997, Photo © Oleg Kosterin

  Larva on Vaccinium [BRU2]: 43, Viola Korb & Bolshakov, 2011, Eversmannia Suppl. 2

  • B. a. angarensis Transbaikalia, S.Siberia

    • Boloria angarensis angarensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana angarensis angarensis; [BRU2], 43; [RFEL]
  • B. a. herzi (Wnukowsky, 1927) N.Transbaikalia, Far East Yakutia

    • Argynnis (Brentis[sic]) angarensis herzi Wnukowsky, 1927; Mitt. Münch. Ent. Ges. 17 (7-12): 70; TL: Jakutsk

    • Clossiana angarensis herzi; [BARU]; [BRU2], 43; [LepIndex]; [CoL]
  • B. a. samkoi (Sheljuzhko, 1931) Polar Urals, Yamal Peninsula

    • Brenthis selenis samkoi Sheljuzhko, 1931; Int. ent. Zs. 25: 45; TL: Dorfe Durynina
    • Brenthis selenis samkoi ab. isabella Sheljuzhko, 1931; Int. ent. Zs. 25: 46

    • Boloria angarensis samkoi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana angarensis samkoi; [BARU]; Yakovlev & Kareev, 2013, Caucasian ent. J. 9 (1): 173, pl. 23, f. 3-6; [LepIndex]; [CoL]
    • Clossiana selenis samkoi; [BRU2], 44
  • B. a. sedychi (Weiss, 1964) Polar Urals, Yamal Peninsula

    • Clossiana angarensis sedychi Weiss, 1964; Reichenbachia 4: 16; TL: Komi

    • Clossiana angarensis sedychi; [BRU2], 43; Yakovlev & Kareev, 2013, Caucasian ent. J. 9 (1): 173 (note); [LepIndex]; [CoL]
  • B. a. alticola (Sushkin & Tshetverikov, 1907) Sayan, Tuva (mountains)

    • Argynnis angarensis alticola Sushkin & Tshetverikov, 1907; Horae Soc. Ent. Rus. 38 (1-2): 18
    • Boloria angarensis shelkovnikovi Korb & Yakovlev, 1997; Alexanor 20 (1): 45; TL: Altai, Oukok, Karaboulak, 2200-2500m
      • = Clossiana angarensis alticola; Yakovlev & Kareev, 2013, Caucasian ent. J. 9 (1): 173

    • Clossiana angarensis alticola; [BARU]; [BRU2], 43; Yakovlev & Kareev, 2013, Caucasian ent. J. 9 (1): 173, pl. 23, f. 9-12; [LepIndex]; [CoL]
  • B. a. miyakei (Matsumura, 1919) Sakhalin

    • Argynnis amathusia var. miyakei Matsumura, 1919; Thous. Ins. Japan. Addit. 3: 584, pl. 45, f. 8-9; TL: Saghalien (Korsarcoff)

    • Argynnis amathusia miyakei; Esaki, 1922, Zool. Mag. Tokyo 34: 907; Matsumura, 1925, J. Coll. Agric. Hokkaido imp. Univ. 15: 99
    • Boloria angarensis miakei[sic, miyakei]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana angarensis miakei[sic, miyakei]; [BARU]; [BRU2], 43
    • Clossiana angarensis miyakei; [RFEL]
    • Clossiana titania miyakei; [BARU]; [LepIndex]; [CoL]
  • B. a. hakutozana (Matsumura, 1927) Amur, Ussuri, N.Korea, NE.China

    • Argynnis hakutozana Matsumura, 1927; Insecta Matsumurana 1 (4): 165, pl. 5, f. 13; TL: Korea

    • Argynnis hakutozana; [LepIndex] ; [CoL]
    • Boloria angarensis hakutozana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana angarensis hakutozana; [BRU2], 43; [RFEL]
    • Clossiana hakutosana[sic, hakutozana]; [EBW], (name only)
    • Clossiana hakutozana; [BARU], #195
• Boloria thore (Hübner, [1803-1804]) Japan. See [maps] Alpen-Perlmutterfalter Gråkantad pärlemorfjäril Bäckpärlefjäril Thore's Fritillary Purohopeatäplä

  • Papilio thore Hübner, [1803-1804]; Samml. eur. Schmett. [1]: pl. 111, f. 571-573; TL: Alps of Tirol
  • Argynnis thore var. jezoensis Matsumura, 1919; Thous. Ins. Japan. Addit. 3: 586; TL: Hokkaido (Kitami)
  • Argynnis thore f. daphnoides Stauder, 1922; Ent. Anz. 2 (4): 46
  • Argynnis thore ab. albina Foltin, 1938; Zs. Öst. EntVer. 23: 125

  • Boloria thore; [BRU2]: 46, pl. 29, f. 22-24; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SVF]
  • Clossiana thore daphnoides; [LepIndex]; [CoL]
  • Clossiana thore jezoensis; Kudrna, 1974, Atalanta 5: 109; [LepIndex]; [CoL]
  • Clossiana thore; [H&R], 107; [BOW]: pl. 2, f. 4; [BARU], #194; [MRS], 478; [RFEL]; [LepIndex] ; [CoL] ; [Mario Maier]

  600x600 (~98Kb) upperside forest-tundra belt of the Koni Peninsula, S Magadan Region, Russia. 11th of July 1989, Photo © Oleg Kosterin	374x506 (~29Kb) underside Among the dead branches of the Siberian dwarf pine (Pinus pumila Ledeb.) A forest/tundra on the Koni Peninsula, S Magadan Region, Russia. 11th of July 1989, Photo © Oleg Kosterin	494x684 (~70Kb) upperside
  470x733 (~69Kb) underside

  Larva on Viola selkirkii, Viola grypoceras [MRS], Viola bifolia, (Finland) Viola altaica, (Urals) [BRU2]: 47

  • B. t. thore Alps

    • Clossiana thore thore; [H&R], 110; [BRU2], 46
  • B. t. borealis (Staudinger, 1861) N.Scandinavia, N.Siberia, Far East, Kamchatka

    • Argynnis thore var. borealis Staudinger, 1861; in Staudinger & Wocke, Cat. Lep. (ed. 1): 9

    • Boloria thore borealis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist]
    • Clossiana thore borealis; [H&R], 110; [BRU2], 47; [RFEL]; [LepIndex]; [CoL]
  • B. t. transuralensis (Sheljuzhko, 1931) Ural, W.Siberia

    • Brenthis thore transuralensis Sheljuzhko, 1931; Int. ent. Zs. 25: 46; TL: "Umgegend von Tobolsk"

    • Boloria thore transuralensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist]
    • Clossiana thore transuralensis; [BARU]; [BRU2], 47; [LepIndex]; [CoL]
  • B. t. hyperlampra (Fruhstorfer, 1907) Pyrenees

    • Argynnis thore hyperlampra Fruhstorfer, 1907; Ent. Zs. 21 (27): 164; TL: Pyrenees

    • Clossiana thore hyperlampra; [LepIndex]; [CoL]
  • B. t. hypercala (Fruhstorfer, 1907) Altai, Sayan, Transbaikalia

    • Argynnis thore hypercala Fruhstorfer, 1907; Ent. Zs. 21 (27): 164
      • = Clossiana thore borealis; [RFEL]
    • Clossiana thore lacrimosa Korb, 1999; Lambillionea 99 (4): 571; TL: Altai

    • Boloria thore hypercala; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana thore hypercala; [LepIndex]; [CoL]

    600x600 (~102Kb) ♀ upperside An open spruce forest, the Stolby Nature Reserve, Krasnoyarsk Province, 11th July 1995, Photo © Oleg Kosterin	784x1098 (~137Kb) ♂ ♀ upperside A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin	800x800 (~145Kb) ♀ underside A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
    1088x786 (~121Kb) ♂ underside A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin

  • B. t. herculea Fruhstorfer, 1907² ?

    • Clossiana thore herculea Fruhstorfer, 1907²
    • herculea Fruhstorfer, 1907; ; TL: Altai

    • Clossiana thore herculea; [BARU]
  • B. t. hyperusia (Fruhstorfer, 1907) Amur, Ussuri

    • Argynnis thore hyperusia Fruhstorfer, 1907; Ent. Zs. 21 (27): 164; TL: Amur
      • = Clossiana thore borealis; [RFEL]
    • Argynnis thore meridionalis Kardakoff, 1928; Ent. Mitt. 17 (4): 270, pl. 7, f. 5-6
      • = Clossiana thore hyperusia; [BRU2], 47
    • Clossiana brenda Hemming, 1933; Entomologist 66 (12): 276 (repl. Argynnis thore meridionalis Kardakoff, 1928)
      • = Clossiana thore hyperusia; [BRU2], 47

    • Boloria thore hyperusia; [iNaturalist]
    • Clossiana thore brenda; [LepIndex]; [CoL]
    • Clossiana thore hyperusia; [BARU]; [MRS], 478; [BRU2], 47; [LepIndex]; [CoL]
    • Clossiana thore meridionalis; [BARU]
  • B. t. excellens (Krulikovsky, 1893)

    • Argynnis thore var. excellens Krulikovsky, 1893; Soc. Ent. 7 (21) (24): 187; TL: Rossia orientalis
    • Clossiana splendida (Krulikovsky, 1895)

    • Argynnis thore excellens; Fruhstorfer, 1907, Ent. Zs. 21 (27): 164
    • Clossiana thore excellens; [BRU2], 47; [LepIndex]; [CoL]
  • B. t. karafutonis (Matsumura, 1919) Sakhalin

    • Argynnis thore var. karafutonis Matsumura, 1919; Thous. Ins. Japan. Addit. 3: 586; TL: Saghalien (Korarkof, Solowiyofka)

    • Clossiana thore karafutonis; [BRU2], 47; [LepIndex]; [CoL]
• Boloria frigga (Thunberg, 1791) . See [maps] Hjortronpärlefjäril Rahkahopeatäplä Frigga's Fritillary Friggas pärlemorfjäril Willow-Bog Fritillary

  • Papilio frigga Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 33 (& Becklin ?); TL: Lappland
  • Argynnis frigga ab. ahti Sahlberg, 1891; Medd. Soc. Fenn. 18: 205, f. 4; TL: Härkäsuo, Sammatti
  • Argynnis frigga ab. tapio Sahlberg, 1891; Medd. Soc. Fenn. 18: 205, f. 5; TL: Härkäsuo, Sammatti
  • Argynnis frigga f. ahti Stichel, 1908; Berl. ent. Z. 53 (1): 86, pl. 3, f. 7
  • Argynnis frigga f. tapio Stichel, 1908; Berl. ent. Z. 53 (1): 86, pl. 3, f. 8
  • Brenthis frigga lapponica Lingonblad, 1944; Notulae Ent. 24 (1-2) (1-2): 51; TL: Finland, Lappland
  • Brenthis (Clossiana) frigga australis Lingonblad, 1946; Notulae Ent. 26 (1-2): 55 (preocc. Argynnis oscarus var. australis Graese, 1888); TL: Finland, Muonio, Enontekiö
  • exmaculata (Roos, 1953) (Clossiana); Opusc. ent. 18: 229

  • Boloria (Clossiana) frigga; Pelham, 2008, J. Res. Lepid. 40: 300; [P&N23]
  • Boloria frigga; [BNA]: 321, #244, pl. 26; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [MPG] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [BOA] ; [BAMONA] ; [SVF]
  • Brenthis frigga; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • Clossiana frigga ahti; [LepIndex]; [CoL] ; [iNaturalist]
  • Clossiana frigga australis; [LepIndex]; [CoL] ; [iNaturalist]
  • Clossiana frigga tapio; [LepIndex]; [CoL] ; [iNaturalist]
  • Clossiana frigga; [SPRK]; [H&R], 110; [NACL], #4466; [BARU], #196; [BOW]: pl. 1, f. 27; [BRU2]: 47, pl. 32, f. 13-23; [RFEL]; [LepIndex] ; [CoL]

  461x381 (~37Kb) upperside

  447x373 (~36Kb) underside

  Larva on Rubus chamaemorus [SPRK], Salix , Betula , Dryas integrifolia [BNA]: 321

  • B. f. frigga N.Scandinavia

    • Boloria frigga helvenacius Sedykh, 1977; Geogr. Aspect. Conserv. Flora Faun. NE Europ. USSR: 102; TL: Polar Urals
      • = Boloria frigga; Pelham, 2008, J. Res. Lepid. 40: 300

    • Boloria frigga frigga; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana frigga frigga; [BRU2], 47
  • B. f. saga (Staudinger, 1861) Canada

    • Argynnis frigga var. saga Staudinger, 1861; Stettin Ent. Ztg 22 (10-12): 350 (Kaden); TL: Labrador

    • Argynnis frigga var. saga; Strecker, 1878, Synon. Cat. Macroplep.: 117
    • Boloria (Clossiana) frigga saga; [P&N23]
    • Boloria frigga saga; [BNA], #244; Pelham, 2008, J. Res. Lepid. 40: 300; [iNaturalist] ; [BOA]
    • Brenthis frigga saga; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
    • Clossiana frigga saga; [NACL], #4466a; [LepIndex]; [CoL]
  • B. f. alpestris (Elwes, 1899) Altai, Sayan, Transbaikalia

    • Argynnis frigga var. alpestris Elwes, 1899; Trans. ent. Soc. Lond. 1899 (3): 340

    • Boloria frigga alpestris; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana frigga alpestris; [BARU]; [BRU2], 47; Yakovlev, 2012, Nota lepid. 35 (1): 90; [LepIndex]; [CoL] ; [iNaturalist]

    872x600 (~56Kb) ♀ upperside Highland dwarf birch tundras on of the southerm principal slope of the Yuzhno-Chuiskii mountain at the the Chikty Rivulet headwaters, 2400 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin	900x592 (~98Kb) ♀ upperside A flowery alpine meadow in a valley of the rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2600 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 11th July 1998, Photo © Oleg Kosterin	800x800 (~121Kb) ♀ underside A flowery alpine meadow in a valley of the rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2600 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 11th July 1998, Photo © Oleg Kosterin
    475x500 (~33Kb) ♂ underside An alpine meadow vegetation at a brook flowing in a moss/sedge tundra in a cirque of the sources of a rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2650 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16 July 1998, Photo © Oleg Kosterin

  • B. f. annae (Suschkin, 1906) Tarbgatai Mts.

    • Argynnis frigga annae Suschkin, 1906; Rev. russ. ent. 6: 5; TL: Tarbagatai

    • Boloria frigga annae; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana frigga annae; [BRU2], 47 (?); [LepIndex]; [CoL] ; [iNaturalist]
  • B. f. sagata (Barnes & Benjamin, 1923) Wyoming - Colorado

    • Brenthis frigga r. sagata Barnes & Benjamin, 1923; Can. Ent. 55 (6): 146; TL: Hall Valley, Colorado

    • Boloria (Clossiana) frigga sagata; [P&N23]
    • Boloria frigga sagata; [BNA], #244; Pelham, 2008, J. Res. Lepid. 40: 300; [iNaturalist] ; [BOA]
    • Clossiana frigga sagata; [NACL], #4466c; [LepIndex]; [CoL]
  • B. f. gibsoni (Barnes & Benjamin, 1926) Alaska - Hudson Bay

    • Brenthis frigga r. gibsoni Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 92; TL: Barter I., Northern Alaska
    • Argynnis frigga f. alaskensis Lehmann, 1913; Gross-Schmett. Erde 5: 424, pl. 87 e (preocc. Brenthis pales var. alaskensis Holland, 1900); TL: NE.Alaska, 69°40N, 141°W
      • = ; Pelham, 2008, J. Res. Lepid. 40: 300
      • = Boloria frigga kosarevi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Brenthis frigga var. lehmanni Holland, 1928; Ann. Carnegie Mus. 19 (1): 44 (repl. Argynnis frigga f. alaskensis Lehmann, 1913); TL: NE.Alaska, 69°40N, 141°W
      • = ; Pelham, 2008, J. Res. Lepid. 40: 301
    • Boloria frigga kosarevi Korb, 2011; Eversmannia Suppl. 2: 33 (repl. Argynnis frigga f. alaskensis Lehman, 1913); TL: NE.Alaska, 69°40N, 141°W
      • = ; Korb & Bolshakov, 2016, Zootaxa 4160 (1): (1-234)

    • Boloria (Boloria) alaskensis; [P&N23]
    • Boloria (Clossiana) frigga gibsoni; [P&N23]
    • Boloria frigga gibsoni; [BNA], #244; Pelham, 2008, J. Res. Lepid. 40: 300; [iNaturalist] ; [BOA]
    • Clossiana frigga alaskensis; [NACL], #4466b
    • Clossiana frigga gibsoni; [LepIndex]; [CoL]
  • B. f. maritima (Kardakoff, 1928) Amur, Ussuri

    • Argynnis frigga maritima Kardakoff, 1928; Ent. Mitt. 17 (4): 270, pl. 7, f. 4; TL: Primorye, Ternei Bay
      • = Boloria frigga; Pelham, 2008, J. Res. Lepid. 40: 300

    • Boloria frigga maritima; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana frigga maritima; [BARU]; [BRU2], 47; [RFEL]; [LepIndex]; [CoL] ; [iNaturalist]
• Boloria dia (Linnaeus, 1767) -. See [maps] Dostojka dia Violet Fritillary Perlada Violeta Hainveilchen-Scheckling La Petite Violette

  • Papilio dia Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 785; TL: Austria
  • Papilio dia minor Borkhausen, 1788; Naturges. Eur. Schmett. 1: 41
  • Brenthis nenoquis Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3): 247; TL: California [error]
    • = Argynnis dia; Strecker, 1878, Synon. Cat. Macroplep.: 118
    • = ; Pelham, 2008, J. Res. Lepid. 40: 442
  • Argynnis dia ab. vittata Spuler, 1901; Schmett. Eur. 1: 28
  • Argynnis dia ab. nigrostriata Schultz, 1908; Ent. Zs. 22 (9): 39
  • Argynnis dia ab. senta Schultz, 1908; Ent. Zs. 22 (10): 40
  • Argynnis dia ab. mediofasciata Schultz, 1908; Soc. Ent. 22 (23): 177
  • Argynnis dia diniensis Oberthür, 1909; Étud. Lépid. Comp. 3: 216
  • Argynnis dia ab. ornata Aigner-Abafi, 1906; Ann. hist.-nat. Mus. Hung. 4 (2): 503
  • Argynnis dia ab. hudaki Aigner-Abafi, 1906; Rovart. Lapok 13 (4): 72
  • Argynnis dia ab. hudaki Aigner-Abafi, 1906; Ent. Zs. 19 (35): 208; TL: Vinkovece
  • Argynnis dia ab. lugens Grund, 1908; Soc. ent. 23 (11): 82
  • nigricans (Ksienschopolski, 1911) (Argynnis); Trudy obshch.Izsl. Volyni Zitomir 8: 40, pl. 1, f. 5
  • Brenthis dia r. laetior Verity, 1919; Ent. Rec. J. Var. 31: 194
  • Brenthis dia f. flavens Verity, 1919; Ent. Rec. J. Var. 31: 194
  • interligata (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 22; TL: France
  • ?euphrosynella (Cabeau, 1924) (Argynnis); Rev. mens. Soc. ent. Namur. 24: 34
  • bivittata (Cabeau, 1925) (Argynnis); Rev. mens. Soc. ent. Namur. 25: 7; TL: Belgium
  • posteronubilata (Cabeau, 1930) (Argynnis); Lambillionea 30: 122
  • Boloria dia f. postdia Verity, 1933; Bull. Soc. ent. Fr. 38 (5): 74
  • Argynnis dia f. albomaculata Bedrnik, 1936; Polsk. Pismo ent. 13: 197
  • couleti (Oberthür, 1937) (Argynnis); Lambillionea 37: 26; TL: Basses-Alpes
  • reducta (Lumma, 1938) (Argynnis); Ent. Zs. 52 (13): 104

  • Argynnis dia diniensis; Fruhstorfer, 1917, Archiv Naturg. 82 A (2): 18
  • Boloria (Clossiana) dia; [P&N23]
  • Boloria dia; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SVF]
  • Boloria euphrosynella; [iNaturalist]
  • Clossiana dia albomaculata; [LepIndex]; [CoL]
  • Clossiana dia; [H&R], 107; [BOW]: pl. 1, f. 28; [EBW]; [BARU], #200; [MRS], 477; [BRU2]: 49, pl. 33, f. 13-21; [RFEL]; [LepIndex] ; [CoL] ; [Mario Maier]
  • Clossiana postdia; Verity, 1950, Le Farfalle diurn. d'Italia 4: 247

  533x400 (~44Kb) underside A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 129 - 15th, 130 - 12th July, 1988, Photo © Oleg Kosterin	598x400 (~44Kb) larva Central Europe 1990-2000, Photo © Michael Borsch	598x400 (~45Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch
  1100x768 (~335Kb) upperside Russia, West Siberia, Altaiskii Krai Province, Zmeinogorsk District, Savvushka village environs, the Lake Kolyvenskoe bank, a slope steppe. 9th September 2007, Photo © Oleg Kosterin	465x500 (~43Kb) upperside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes	500x503 (~50Kb) underside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes

  Larva on Viola , Rubus idaeus, Prunella vulgaris [BRU2]: 49

  • B. d. dia W.Europe

    • Boloria dia dia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist]
    • Clossiana dia dia; [BRU2], 49
  • B. d. nigrescens (Schavrov, 1886)

    • dia nigrescens (Schavrov, 1886) (Argynnis); ? 50: 203

    • Boloria dia nigrescens; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
  • B. d. leonina (Fruhstorfer, 1909)

    • Argynnis dia leonina Fruhstorfer, 1909; Int. ent. Zs. 3 (4): 21; TL: Savoyen

    • Clossiana dia leonina; [LepIndex]; [CoL]
  • B. d. setania (Fruhstorfer, 1909)

    • Argynnis dia setania Fruhstorfer, 1909; Int. ent. Zs. 3 (4): 21; TL: 'Marokko' [mislabeling?]

    • Clossiana dia setania; [BMAT]: 105, pl. 32, f. 19-20 (not in N.Africa); [LepIndex]; [CoL]
  • B. d. disconota (Krulikovsky, 1909) CEU, W.Siberia

    • Argynnis dia var. disconota Krulikovsky, 1909; Rev. russ. ent. 9 (3): 298

    • Clossiana dia disconota; [BRU2], 49
  • B. d. calida (Jachontov, 1911) Caucasus, Transcaucasus

    • Argynnis dia var. calida Jachontov, 1911; Mitt. Kaukas. Mus. 5: 306

    • Clossiana dia calida; [BRU2], 49; [LepIndex]; [CoL]
  • B. d. alpina (Elwes, 1899) Dzhungarsky Alatau, Altai, Sayan, Transbaikalia

    • Argynnis dia var. alpina Elwes, 1899; Trans. ent. Soc. Lond. 1899 (3): 339; TL: Altai

    • Boloria dia alpina; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist]
    • Clossiana dia alpina; [BARU]; [BRU2], 49; Yakovlev, 2012, Nota lepid. 35 (1): 90; [LepIndex]; [CoL]
  • B. d. semota (Tuzov, 1999) Magadan

    • Clossiana dia semota Tuzov, 1999; Helios 1: 105; TL: Russia, Magadan

    • Boloria dia semota; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana dia semota; [BRU2], 49; [RFEL]
• Boloria gong (Oberthür, 1884) Tibet, W.China, N.China. See [maps]

  • Argynnis gong Oberthür, 1884; Étud. d'Ent. 9: 15, pl. 2, f. 9; TL: Ta-Tsien-Lou
  • Argynnis eva Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 456
  • Argynnis gong pernimia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 17; TL: China, Kansu
  • Argynnis gong f. pallida Eisner, 1942; Zool. Meded. Leiden 24 (4): 122; TL: Sung-pan-ting, Szetschwan

  • Boloria gong; [iNaturalist]
  • Clossiana eva; [LepIndex] ; [CoL]
  • Clossiana gong pallida; [LepIndex]; [CoL]
  • Clossiana gong pernimia; [LepIndex]; [CoL]
  • Clossiana gong; [BOW]: pl. 194, f. 11; [MRS], 477; Coene & Vis, 2008, Nota lepid. 1 (2): 239; [LepIndex] ; [CoL]
  • B. g. charis (Oberthür, 1891) Yunnan, SE.Tibet

    • Argynnis charis Oberthür, 1891; Étud. d'Ent. 15: 8, pl. 1, f. 4; TL: Yunnan
    • Clossiana gong xizangensis Huang, 2000
      • = Clossiana gong charis; Coene & Vis, 2008, Nota lepid. 1 (2): 239

    • Clossiana eva charis; [LepIndex]; [CoL]
    • Clossiana gong charis; Coene & Vis, 2008, Nota lepid. 1 (2): 239
  • ?B. g. splendens (Murayama, 1983)

    • Clossiana gong splendens Murayama, 1983; Entomotaxonomia 5: (281-288)

    • ?Clossiana gong splendens; [MRS], 477
• Boloria titania (Esper, 1794) . See [maps] Amathusias pärlemofjäril Lundpärlefjäril Natterwurz-Perlmutterfalter Purple Bog Fritillary Titania's Fritillary Dostojka titania Lehtohopeatäplä

  • Papilio titania Esper, 1794; Die Schmett., Suppl. Th 1 (3-4): 58, (5-6): pl. 103, f. 4; TL: Sardinia
  • Papilio dia major Esper, 1786; Die Schmett. Th. I, Bd. 2 (28): pl. 93, f. 2-3
  • Papilio diana Hübner, [1799-1800]; Samml. eur. Schmett. [1]: pl. 10, f. 51-54
  • Argynnis amathusia ab. radiifera Schultz, 1908; Soc. Ent. 22 (23): 177
  • Argynnis amathusia ab. nigrata Schultz, 1908; Soc. Ent. 22 (23): 177
  • Argynnis amathusia var. transsylvanica Tiltscher, 1913; Ent. Zs. 26: 210
  • Argynnis amathusia ab. bialoviezensis Gieysztor, 1923; Bialowieza 2: 125
  • Boloria amathusia titania f. microtitania Verity, 1933; Ent. Rec. J. Var. 45 (6): 91
  • Argynnis amathusia nansetsuzana Doi, 1935; Zephyrus 6: 15; TL: Korea
  • lemagneni (Plantrou, 1960) (Clossiana); Alexanor 1: 199

  • Boloria titania; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [iNaturalist] ; [LEPIFORUM] ; [Heiner Ziegler] ; [MBE] ; [MBE] ; [SVF]
  • Clossiana titania lemagneni; [LepIndex]; [CoL]
  • Clossiana titania nansetsuzana; [LepIndex]; [CoL]
  • Clossiana titania transsylvanica; [LepIndex]; [CoL]
  • Clossiana titania; [H&R], 105; [BOW]: pl. 2, f. 5; [EBW]; [BARU], #198; [BRU2]: 49, pl. 33, f. 31-36; [RFEL]; [LepIndex] ; [CoL] ; [Mario Maier]

  Misidentified/misapplied:

  • Clossiana titania (= Clossiana chariclea); [NACL], #4474
  • Boloria titania (= Clossiana chariclea); [BNA]: 319, #240

  571x653 (~100Kb) Russia, Moscow area (36°25'E, 56°00'N), 24.7.2009, Photo © D. Smirnov

  604x640 (~97Kb) underside Russia, Moscow area (36°25'E, 56°00'N), 24.7.2009, Photo © D. Smirnov

  Larva on Polygonum , Filipendula , Trollius , Viola [RFEL]

  • B. t. titania Sardinia, ?

    • Clossiana titania titania; [H&R], 105

    Larva on Viola , Polygonum [H&R]

  • B. t. cypris (Meigen, 1829) S.Germany, Switzerland

    • titania cypris (Meigen, 1829) (Argynnis); Syst. Beschr. eur. Schmett. 2: 5

    • Boloria titania cypris; Korb & Bolshakov, 2011, Eversmannia Suppl. 2:
    • Clossiana titania cypris; [H&R], 106
  • B. t. bivina (Fruhstorfer, 1908) CEU, SEU

    • Argynnis amathusia bivina Fruhstorfer, 1908; Int. ent. Zs. 1 (41): 310; TL: Saratow
    • Argynnis amathusia bivina f. blandina Fruhstorfer, 1908; Int. ent. Zs. 1 (41): 310; TL: Zermatt
    • Papilio amathusia Esper, 1783; Die Schmett. Th. I, Bd. 2 (8): 170, pl. 88, f. 1-2 (preocc.)
    • rossicus (Hemming, 1933) (Argynnis); Entomologist 66: 276 (repl. Papilio amathusia Esper, 1783)

    • Clossiana titania bivina; [BARU]; [BRU2], 50; [LepIndex]; [CoL]
    • Clossiana titania blandina; [LepIndex]; [CoL]
    • Clossiana titania rossicus; [LepIndex]; [CoL]

    Larva on Viola , Vaccinium uliginosum [BRU2]: 50

  • B. t. bosna (Fruhstorfer, 1908) Bosnia

    • Argynnis amathusia bosna Fruhstorfer, 1908; Int. ent. Zs. 2 (33): 214; TL: Bosnia, Trebevic

    • Clossiana titania bosna; [LepIndex]; [CoL]
  • B. t. dinara (Fruhstorfer, 1908) Herzegowina

    • Argynnis amathusia dinara Fruhstorfer, 1908; Int. ent. Zs. 2 (33): 214; TL: Herzegovina, Gacko; Vucija-bara

    • Clossiana titania dinara; [LepIndex]; [CoL]
  • B. t. serena (Fruhstorfer, 1908) S.Tirol

    • Argynnis amathusia serena Fruhstorfer, 1908; Int. ent. Zs. 2 (33): 214; TL: Wallis, Zermatt and Simplon region, S.Tirol, Piemont, Courmayeur

    • Clossiana titania serena; [LepIndex]; [CoL]
  • B. t. pralognana (Fruhstorfer, 1908)

    • Argynnis amathusia pralognana Fruhstorfer, 1908; Int. ent. Zs. 2 (33): 214; TL: Pralognan, ca. 1400-1500m

    • Clossiana titania pralognana; [LepIndex]; [CoL]
  • B. t. halesa (Fruhstorfer, 1908) Pyrenees

    • Argynnis amathusia halesa Fruhstorfer, 1908; Int. ent. Zs. 2 (33): 214; TL: Pyrenees

    • Clossiana titania halesa; [LepIndex]; [CoL]
  • B. t. blachieri (Fruhstorfer, 1909)

    • Argynnis amathusia blachieri Fruhstorfer, 1909; Int. ent. Zs. 3 (4): 20; TL: Tessin, Fusio

    • Clossiana titania blachieri; [LepIndex]; [CoL]
  • B. t. jugurtha (Fruhtorfer, 1909)

    • Argynnis amathusia jugurtha Fruhstorfer, 1909; Int. ent. Zs. 3 (4): 20; TL: Baeyern, Kochelsee, Berchesgaed, Triol, Karrersee

    • Clossiana titania jugurtha; [LepIndex]; [CoL]
  • B. t. staudingeri (Wnukowsky, 1929) S.Siberia, Altai, Sayan, mountains (Tuva), Transbaikalia, Amur, Sakhalin?

    • titania staudingeri (Wnukowsky, 1929) (Brenthis); Zool. Anz. 83 (9/10): 222 (repl. Argynnis amathusia var. sibirica Staudinger, 1892); TL: Kentei
    • Argynnis amathusia var. sibirica Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 330 (preocc. Argynnis selenis var. sibirica Erschoff, 1870); TL: Kentei
    • Argynnis amathusia f. altaica Seitz, 1909; Gross-Schmett. Erde 1: 232, pl. 68 f (preocc. Argynnis pales var. altaica Grum-Grshimailo, 1893); TL: Kentei
    • Argynnis titania tana Hemming, 1934; Stylops 3 (9): 193 (repl. Argynnis amathusia f. altaica Seitz, 1909)
    • Argynnis titania julia Hemming, 1934; Stylops 3 (5): 97 (repl. Argynnis amathusia f. altaica Seitz, 1909)

    • Boloria titania staudingeri; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana titania julia; [LepIndex]; [CoL]
    • Clossiana titania staudingeri; [BARU]; [BRU2], 50; Yakovlev, 2012, Nota lepid. 35 (1): 91; [RFEL]; [LepIndex]; [CoL]

    700x510 (~58Kb) upperside A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin

    Larva on Bistorta major, Filipendula ulmaria, Trollius asiaticus [BRU2]: 50

  • B. t. minuscula (Eisner, 1942) Lithuania

    • Argynnis amathusia minuscula Eisner, 1942; Zool. Meded. Leiden 24 (4): 122; TL: Amata, Lettland

    • Clossiana titania minuscula; [LepIndex]; [CoL]
• Boloria chariclea (Schneider, 1794) . See [maps] Tundrapärlefjäril Arctic Fritillary Tundrahopeatäplä

  • Papilio chariclea Schneider, 1794; Neuest. Mag. Lieb. Ent. 1 (5): 588; TL: Lappland
  • aili (Kontuniemi, 1939) (Argynnis); Ann. ent. fenn. 5: 165

  • Boloria (Clossiana) chariclea; Pelham, 2008, J. Res. Lepid. 40: 305; [P&N23]
  • Boloria chariclea; [BOC], 286; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [MPG] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [BOA] ; [BAMONA] ; [SVF]
  • Brenthis chariclea; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • Clossiana carichlea; Zetterstedt, 1839, (missp.)
  • Clossiana chariclea; [H&R], 106; [NACL], #4475; [BARU], #199; [BOW]: pl. 1, f. 25; [BRU2]: 48, pl. 33, f. 4-9; [RFEL]; [LepIndex] ; [CoL] ; [SZMN]
  • Clossiana churchilea; dos Passos, 1964, (missp.)
  • Clossiana churiclea; Johansen, 1921, (missp.)

  550x424 (~48Kb) upperside FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela

  500x564 (~35Kb) underside FINLAND: Li: Utsjoki Ailigas, 775:50, 8.7.1997, Photo © Markku Savela

  700x506 (~69Kb) upperside FINLAND: Li: Utsjoki Ailigas, 775:50, 6.7.1997, Photo © Markku Savela

  Larva on Dryas integrifolia, Salix arctica, S. reticulata, (N.America) [BRU2]: 49

  • B. c. chariclea Arctic Europe

    • Boloria chariclea chariclea; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana chariclea chariclea; [BRU2], 48

    Larva on Cassiope tetragona [BRU2]: 49

  • B. c. boisduvalii (Duponchel, 1832) Alaska, Alberta, Labrador, Newfoundland, Minnesota, British Columbia

    • Argynnis boisduvalii Duponchel, 1832; in Godart, Hist. nat. Lépid. Fr. (Suppl.) 1: pl. 20, f. 4
    • Argynnis oenone Scudder, 1875; Proc. Boston Soc. nat. Hist. 17 (3): 299
      • = ; Pelham, 2008, J. Res. Lepid. 40: 305 (nom. nud.)
    • Brenthis chariclea r. oenone tr.f. labradorensis Gunder, 1928; Can. Ent. 60 (7): 163, pl. A, f. 5, 5a; TL: Labrador
      • = ; Pelham, 2008, J. Res. Lepid. 40: 305

    • Boloria (Clossiana) chariclea boisduvalii; [P&N23]
    • Boloria chariclea boisduvalii; Pelham, 2008, J. Res. Lepid. 40: 305; [BOA]
    • Boloria titania boisduvalii; [BNA], #240
    • Brenthis chariclea boisduvalii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
    • Clossiana boisduvalii; [LepIndex] ; [CoL]
    • Clossiana titania boisduvalii; [NACL], #4474a
  • B. c. arctica (Zetterstedt, 1839) Arctic Asia, Wrangel Is., Chukotka

    • Argynnis arctica Zetterstedt, 1839; Insecta Lapponica: 899
    • Papilio tullia Fabricius, 1780; Fauna Groenl.: 192 (preocc. Papilio tullia Müller, 1764)
      • = ; Pelham, 2008, J. Res. Lepid. 40: 305
    • Argynnis chariclea var. obscurata McLachlan, 1878; J. Linn. Soc. Zool. Lond. 14: 110
      • = ; Pelham, 2008, J. Res. Lepid. 40: 306

    • Argynnis chariclea var. obscurata; Holland & Avinoff, 1935, Mem. Carn. Mus. 12 (2): 4
    • Boloria (Clossiana) chariclea arctica; [P&N23]
    • Boloria chariclea arctica; Pelham, 2008, J. Res. Lepid. 40: 305; [iNaturalist] ; [BOA]
    • Brenthis chariclea arctica; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
    • Brenthis chariclea obscurata; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
    • Clossiana chariclea arctica; [NACL], #4475a; [LepIndex]; [CoL]
    • Clossiana chariclea obscurata; [LepIndex]; [CoL]
  • B. c. montinus (Scudder, 1863) New Hamspshire, ...

    • Argynnis montinus Scudder, 1863; Proc. Essex Inst. 3: 166; TL: White Mtns, New Hampshire

    • Boloria (Clossiana) chariclea montinus; [P&N23]
    • Boloria chariclea montinus; Pelham, 2008, J. Res. Lepid. 40: 305; [iNaturalist] ; [BOA]
    • Brenthis montinus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16; dos Passos, 1936, Can. Ent. 68: 239-241
    • Clossiana montinus; [LepIndex] ; [CoL]
    • Clossiana titania montina; [NACL], #4474d
  • B. c. helena (Edwards, 1871) Rocky Mountains

    • Argynnis helena Edwards, 1871; Trans. amer. ent. Soc. 3: 268; TL: Mosquito Pass, Park/Lake cos, CO
    • Boloria titania sangredecristo Scott, 1981; Papilio (n.s.) 1: 5; TL: Hermit Pass, Sangre de Cristo Mts, Custer Co., Colorado
      • = ; Pelham, 2008, J. Res. Lepid. 40: 306

    • Boloria (Clossiana) chariclea helena; [P&N23]
    • Boloria chariclea helena; [BOC], 286; Pelham, 2008, J. Res. Lepid. 40: 306; [iNaturalist] ; [BOA]
    • Boloria titania helena; [BNA], #240
    • Brenthis helena; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
    • Clossiana helena; [LepIndex] ; [CoL]
    • Clossiana titania helena; [NACL], #4474f

    802x949 (~107Kb) ♀ upperside USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela

    800x998 (~123Kb) underside USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela

    800x608 (~84Kb) underside USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela

  • B. c. butleri (Edwards, 1883) Arctic America

    • Argynnis butleri Edwards, 1883; Can. Ent. 15 (2): 32; TL: "Cape Thompson, Kotzebue Sound, W.Alaska"

    • Argynnis butleri; [LepIndex] ; [CoL]
    • Boloria (Clossiana) chariclea butleri; [P&N23]
    • Boloria chariclea butleri; Pelham, 2008, J. Res. Lepid. 40: 306; [iNaturalist] ; [BOA]
    • Clossiana butleri; [BRU2]: 49, pl. 33, f. 10-12
    • Clossiana butlerii; Edwards, 1884, (missp.)
    • Clossiana chariclea butleri; [BARU]; [NACL], #4475b
  • B. c. rainieri (Barnes & McDunnough, 1913) Washington

    • Brenthis chariclea rainieri Barnes & McDunnough, 1913; Contr. nat. Hist. Lep. N. Am. 2 (3): 96, pl. 2, f. 1-4; TL: Paradise Valley, Mt. Ra[i]nier, 6-7000ft, [Pierce or Lewis Co.] Wash[ington]

    • Boloria (Clossiana) chariclea rainieri; [P&N23]
    • Boloria chariclea rainieri; [BOC], 286; Pelham, 2008, J. Res. Lepid. 40: 306; [iNaturalist] ; [BOA]
    • Brenthis chariclea ranieri[sic, rainieri]; Barnes & McDunnough, 1913, Contr. nat. Hist. Lep. N. Am. 2 (3): pl. 2, f. 1-4 (orig. missp.)
    • Clossiana chariclea rainieri; [LepIndex]; [CoL]
    • Clossiana titania rainieri; [NACL], #4474b
    • ranieri[sic, rainieri]; Guppy & Kondla, 2002, J. Lep. Soc. 56 (3): 180
  • B. c. grandis (Barnes & McDunnough, 1916) N.British Columbia, Ontario

    • Brenthis chariclea grandis Barnes & McDunnough, 1916; Can. Ent. 48 (7): 223; TL: Hymers, Ontario

    • Boloria (Clossiana) chariclea grandis; [P&N23]
    • Boloria chariclea grandis; [BOC], 286; Pelham, 2008, J. Res. Lepid. 40: 306; [iNaturalist] ; [BOA]
    • Clossiana chariclea grandis; [LepIndex]; [CoL]
    • Clossiana titania grandis; [NACL], #4474c
  • B. c. ingens (Barnes & McDunnough, 1918) Wyoming

    • Brenthis helena ingens Barnes & McDunnough, 1918; Contr. nat. Hist. Lepid. N. Am. 4 (2): 71, 180, pl. 11, f. 5-6; TL: Yellowstone Park
    • Brenthis helena ingens tr.f. martini Gunder, 1934; Can. Ent. 66 (6): 127; TL: Teton trail, Teton Mts, Teton Co., Wyoming
      • = ; Pelham, 2008, J. Res. Lepid. 40: 306

    • Boloria (Clossiana) chariclea ingens; [P&N23]
    • Boloria chariclea ingens; Pelham, 2008, J. Res. Lepid. 40: 306; [iNaturalist] ; [BOA]
    • Clossiana helena ingens; [LepIndex]; [CoL]
    • Clossiana titania ingens; [NACL], #4474e
  • B. c. tshuktsha (Korshunov & Dubatolov, 1998) Chukotka, Kamchatka

    • Clossiana chariclea tshuktsha Korshunov & Dubatolov, 1998; New descr. specif. Butts. Asiatic Russia: 18 [21?]
      • = Boloria chariclea butleri; Pelham, 2008, J. Res. Lepid. 40: 306

    • Boloria chariclea tshuktsha; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana chariclea tshuktsha; [RFEL]; [SZMN]
• Boloria bellona (Fabricius, 1775) Rocky Mountains, Canada, Washington, Oregon. See [maps] Meadow Fritillary

  • Papilio bellona Fabricius, 1775; Syst. Ent.: 517

  • Boloria (Clossiana) bellona bellona; [P&N23]
  • Boloria (Clossiana) bellona; Pelham, 2008, J. Res. Lepid. 40: 299; [P&N23]
  • Boloria bellona; [BNA]: 320, #243, pl. 26; [BOC], 282; [Opler]; [MPG] ; [iNaturalist] ; [BOA] ; [BOA] ; [BAMONA]
  • Brenthis bellona; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • Clossiana bellona; [NACL], #4465; [NAB], 560; [BOW]: pl. 13, f. 19

  Larva on Viola sororia, V. pallens [BNA]: 320

  • B. b. bellona

    • Argynnis bellona fasciata Cockerell, 1889; Entomologist 22 (311): 99
      • = ; Pelham, 2008, J. Res. Lepid. 40: 300
    • Brenthis bellona ab. kleenei Watson, 1921; J. N.Y. Ent. Soc. 29: 171; TL: West Hartford, Connecticut
      • = ; Pelham, 2008, J. Res. Lepid. 40: 300
    • Brenthis bellona ab. pardopsis Holland, 1928; Ann. Carnegie Mus. 19 (1): 45; TL: Crafton, Allegheny Co., Pennsylvania
      • = ; Pelham, 2008, J. Res. Lepid. 40: 300
    • Argynnis ammiralis Hemming, 1933; Entomologist 66 (12): 276 (repl. Papilio bellona Fabricius, 1775)
      • = ; Pelham, 2008, J. Res. Lepid. 40: 300

    • Boloria bellona bellona; [BNA], #243; Pelham, 2008, J. Res. Lepid. 40: 299; [iNaturalist] ; [BOA]
    • Clossiana toddi ammiralis; [LepIndex]; [CoL]
  • B. b. toddi (Holland, 1928) Labrador, N.Manitoba, Northwest Territories

    • Brenthis bellona toddi Holland, 1928; Ann. Carnegie Mus. 19 (1): 45; TL: St. Margaret's River, Quebec

    • Boloria (Clossiana) bellona toddi; [P&N23]
    • Boloria bellona toddi; [iNaturalist] ; [BOA]
    • Boloria belona toddi; [BNA], #243; Pelham, 2008, J. Res. Lepid. 40: 300
    • Clossiana bellona toddi; [NACL], #4465a
    • Clossiana toddi; [LepIndex] ; [CoL]
  • B. b. jenistai Stallings & Turner, 1947

    • Boloria toddi jenistai Stallings & Turner, 1947; Can. Ent. 78 (7-8): 135; TL: Lloydminster, Sask.

    • Boloria (Clossiana) bellona jenistai; [P&N23]
    • Boloria bellona jenistae; [iNaturalist]
    • Boloria bellona jenistai; Pelham, 2008, J. Res. Lepid. 40: 300; [BOA]
    • Clossiana bellona jenistae[sic, jenistai]; [NACL], #4465b
    • Clossiana toddi jenistai; [LepIndex]; [CoL]
• Boloria epithore (Edwards, 1864) Yukon - California. See [maps] Pacific Fritillary Western Meadow Fritillary

  • Argynnis epithore Edwards, 1864; Proc. ent. Soc. Philad. 2 (4): 504 (Boisduval i.l.); TL: Saratoga, Santa Clara, California
  • Argynnis epithore Boisduval, 1869; Ann. Soc. ent. Belg. 12: 58
    • = Boloria epithore epithore; Pelham, 2008, J. Res. Lepid. 40: 301

  • Boloria (Clossiana) epithore epithore; [P&N23]
  • Boloria (Clossiana) epithore; Pelham, 2008, J. Res. Lepid. 40: 301; [P&N23]
  • Boloria epithore; [BNA]: 320, #242, pl. 26; [BOC], 283; [Opler]; [MPG] ; [LepIndex] ; [CoL] ; [iNaturalist] ; [LEPIFORUM] ; [BOA] ; [BOA] ; [BAMONA]
  • Brenthis epithore Boisduval, 1869; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • Clossiana epithore; [NACL], #4469; [NAB], 564

  identification uncertain 600x539 (~44Kb) upperside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela	identification uncertain 400x369 (~20Kb) underside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela	1000x860 (~111Kb) ♀ upperside USA: Washington, Kittitas Co., Tucquala Lake, 24.7.2011, Photo © Markku Savela
  1000x837 (~96Kb) underside USA: Washington, Kittitas Co., Tucquala Lake, 24.7.2011, Photo © Markku Savela	1400x1046 (~252Kb) upperside USA: Washington, Skagit Co., on Sauk Mountain Rd (48°31'23N 121°36'22W), 15.8.2012, Photo © Markku Savela	1000x925 (~153Kb) underside USA: Washington, Skagit Co., on Sauk Mountain Rd (48°31'23N 121°36'22W), 15.8.2012, Photo © Markku Savela
  1000x1249 (~153Kb) USA: Washington, Skagit Co., Sauk Mountain Trail Head & Trail, 15.8.2012, Photo © Markku Savela	1200x957 (~152Kb) upperside USA: Washington, Skagit Co., Sauk Mountain Trail Head & Trail, 15.8.2012, Photo © Markku Savela

  Larva on Viola sempervirens, V. glabella, V. ocellata, V. adunca, V. nephrophylla? [BOC]: 283

  • B. e. epithore California

    • Boloria epithore epithore; [BNA], #242; Pelham, 2008, J. Res. Lepid. 40: 301; [BOA]
  • B. e. chermocki Perkins & Perkins, 1966 Oregon

    • Boloria epithore chermocki Perkins & Perkins, 1966; J. Lep. Soc. 20 (2): 109; TL: 2.9 mi E of Dolph, Yamhill Co., Oregon
    • Boloria epithore uslui Koçak, 1984; Priamus 3 (3): 96 (repl. Boloria epithore borealis Perkins, 1973); TL: Shingle Creek Rd., Keremeos, BC
      • = ; Pelham, 2008, J. Res. Lepid. 40: 302
    • Boloria epithore borealis Perkins, 1973; Bull. Allyn Mus. 11: 6, pl. 1, f. 13-16 (preocc. Argynnis thore borealis Staudinger, 1861); TL: Shingle Creek Rd., Keremeos, BC
      • = ; Pelham, 2008, J. Res. Lepid. 40: 302
    • Brenthis epithore ab. ♀ obscuripennis Gunder, 1926; Ent. News 37 (1): 7; TL: Chilcolin, Br. Columbia
      • = ; Pelham, 2008, J. Res. Lepid. 40: 302

    • Boloria (Clossiana) epithore chermocki; [P&N23]
    • Boloria epithore chermocki; [BNA], #242; Pelham, 2008, J. Res. Lepid. 40: 302; [LepIndex]; [CoL] ; [iNaturalist] ; [BOA]
    • Clossiana epithore borealis; [NACL], #4469b
    • Clossiana epithore chermocki; [NACL], #4469a
  • B. e. sierra Perkins, 1973 California

    • Boloria epithore sierra Perkins, 1973; Bull. Allyn Mus. 11: 10, pl. 1, f. 1-3; TL: Sentinel Dome, Yosemite Nat. Park, California
    • Brenthis epithore ab. eldorado Strand, 1915; Arch. Naturgesch. 80 A (11): 156; TL: Plumas Co. ... Eldorado Co., California
      • = ; Pelham, 2008, J. Res. Lepid. 40: 302
    • Brenthis epithore ab. wawonae Gunder, 1924; Ent. News 35 (5): 156; TL: Wawona, Mariposa Co., California
      • = ; Pelham, 2008, J. Res. Lepid. 40: 302

    • Boloria (Clossiana) epithore sierra; [P&N23]
    • Boloria epithore sierra; Pelham, 2008, J. Res. Lepid. 40: 302; [iNaturalist] ; [BOA]
    • Clossiana epithore sierra; [NACL], #4469c
  • B. e. sigridae (Shepard, 2001) British Columbia

    • Clossiana epithore sigridae Shepard, 2001; Butts. B.C.: 289; TL: BC, St. Elias Mts, Tats Lake, 770m

    • Boloria (Clossiana) epithore sigridae; [P&N23]
    • Boloria epithore sigridae; Pelham, 2008, J. Res. Lepid. 40: 302; [iNaturalist] ; [BOA]
• Boloria improba (Butler, 1877) . See [maps] Dvärgpärlefjäril Dusky-windged Fritillary Dinky Fritillary Kääpiöhopeatäplä Dingy Arctic Fritillary

  • Argynnis improba Butler, 1877; Ent. mon. Mag. 13: 206; TL: "Winter Cove and Cambridge Bay" [NW Territories, Canada]

  • Boloria (Clossiana) improba improba; [P&N23]
  • Boloria (Clossiana) improba; Pelham, 2008, J. Res. Lepid. 40: 301; [P&N23]
  • Boloria improba; [BNA]: 321, #245, pl. 5, 26, 60; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33; [MPG] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [BOA] ; [BAMONA] ; [SVF]
  • Clossiana improba; [NACL], #4467; [H&R], 111; [BARU], #197; [BOW]: pl. 13, f. 20; [BRU2]: 47, pl. 33, f. 22-24; [RFEL]; [LepIndex] ; [CoL]

  600x450 (~38Kb) Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen	800x583 (~75Kb) underside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen	800x549 (~56Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
  800x709 (~76Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen

  Larva on Polygonum viviparum, Salix [BARU], Bistorta vivipara, Salix herbacea [BRU2]: 47, Salix , S. reticulata?, (NW.Terr.) [BNA]: 221

  • B. i. improba Arctic Europe, Arctic Asia

    • Boloria improba improba; Pelham, 2008, J. Res. Lepid. 40: 301; [iNaturalist] ; [BOA]
    • Brenthis frigga improba; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
    • Clossiana improba improba; [BRU2], 47
  • B. i. youngi (Holland, 1900) Alaska, E.Chukotka?

    • Brenthis youngi Holland, 1900; Ent. News 11: 283; TL: Alaska

    • Boloria (Clossiana) improba youngi; [P&N23]
    • Boloria improba youngi; Pelham, 2008, J. Res. Lepid. 40: 301; [iNaturalist] ; [BOA]
    • Brenthis youngi; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
    • Clossiana improba youngi; [NACL], #4467a
    • Clossiana youngi; [LepIndex] ; [CoL]
  • B. i. improbula (Bryk, 1920) N.Findand, N.Sweden

    • Argynnis improba improbula Bryk, 1920; Ent. Tidskr. 41: 131; TL: N.Sweden
    • Brenthis improba improbula ab. elegans Bruun & Schantz, 1950; Notul. Ent. 29 (3-4): 88, pl. 1, f. 11; TL: Finland
    • Brenthis improba improbula ab. nigrofasciata Bruun & Schantz, 1950; Notul. Ent. 29 (3-4): 88, pl. 1, f. 6; TL: Finland
    • Brenthis improba improbula ab. obscura Bruun & Schantz, 1950; Notul. Ent. 29 (3-4): 88, pl. 1, f. 8; TL: Finland
    • Brenthis improba improbula ab. obsoleta Bruun & Schantz, 1950; Notul. Ent. 29 (3-4): 88, pl. 1, f. 9-10; TL: Finland
    • Brenthis improba improbula ab. transversata Bruun & Schantz, 1950; Notul. Ent. 29 (3-4): 88, pl. 1, f. 7; TL: Finland

    • Boloria improba improbula; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
    • Clossiana improba improbula; [RFEL]; [LepIndex]; [CoL]
  • B. i. acrocnema Gall & Sperling, 1980 Colorado (San Juan Mts.) Uncompaghre Fritillary

    • Boloria (Clossiana) acrocnema Gall & Sperling, 1980; J. Lep. Soc. 34 (2): 231; TL: near Uncompahgre Peak, San Juan Mts., Hinsdale Co., Colorado

    • Boloria (Clossiana) improba acrocnema; [P&N23]
    • Boloria acrocnema; [LepIndex] ; [CoL] ; [iNaturalist]
    • Boloria improba acrocnema; [BNA], #245; Pelham, 2008, J. Res. Lepid. 40: 301; [iNaturalist] ; [BOA]

    Larva on Salix nivalis [BNA]

  • B. i. harryi (Ferris, 1984) Wyoming (Wind River Mts.)

    • Clossiana improba harryi Ferris, 1984; Bull. Allyn Mus. 89: 2; TL: nr summit of Bear Ears Trail, 0.5mi NW of Mt. Chauvenet, 3583,. Shoshone Nat. Forest, Fremont Co., Wyoming

    • Boloria (Clossiana) improba harryi; [P&N23]
    • Boloria improba harryi; [BNA], #245; Pelham, 2008, J. Res. Lepid. 40: 301; [iNaturalist] ; [BOA]

    Larva on Salix arctica [BNA]: 321

  • B. i. nunatak Scott, 1998 Alberta

    • Boloria improba nunatak Scott, 1998; Papilio (n.s.) 11: 3; TL: Torrens River, Alberta

    • Boloria (Clossiana) improba nunatak; [P&N23]
    • Boloria improba nunatak; Pelham, 2008, J. Res. Lepid. 40: 301; [BOA]
• Boloria erda (Christoph, 1893) -. See [maps]

  • Argynnis erda Christoph, 1893; Dt. Ent. Z. Iris 6 (1): 87; TL: "Vilui" [Vilui R., Yakutia]

  • Boloria erda; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34; [iNaturalist]
  • Clossiana erda; [BARU], #203; [BRU2]: 50, pl. 35, f. 4-9; [RFEL]; [LepIndex] ; [CoL]

  Larva on Vaccinium vitis-idaea [BRU2]: 50

  • B. e. erda Yakutia

    • Boloria erda erda; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana erda erda; [BRU2], 50; [RFEL]
  • B. e. dulkeiti (Kurentzov, 1970) Magadan

    • Brenthis dulkeiti Kurentzov, 1970; Butterflies of the USSR Far East: 83, 108; TL: Omsukchansky Mts., Magadan
      • = Boloria erda; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34

    • Clossiana erda dulkeiti; [BARU]; [BRU2], 50 (?)
  • B. e. kitoica (Belik, 1996) E.Sayan

    • Clossiana erda kitoica Belik, 1996; Phegea 24 (4): 163; TL: Vostochnyy Sayan mtns

    • Boloria erda kitoica; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana erda kitoica; [BRU2], 50
  • B. e. puella (Churkin, 1999) Transbaikalia (Barguzinsky Mts.)

    • Clossiana erda puella Churkin, 1999; Atalanta 29 (1/4): 121; TL: Buryatia, Barguzin Mts., Nesterikha River, Kedrovoe Lake, 1900-2000m

    • Boloria erda puella; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana erda puella; [BRU2], 50
• Boloria alberta (Edwards, 1890) Alberta, British Columbia. See [maps] Alberta Fritillary Alberta Alpine Fritillary

  • Argynnis alberta Edwards, 1890; Can. Ent. 22 (6): 113; TL: Laggan, Alberta, Canada
  • Brenthis alberta tr.f. banffensis Gunder, 1932; Pan-Pacific Ent. 8 (3): 125; TL: Banff, Alberta, Canada
    • = ; Pelham, 2008, J. Res. Lepid. 40: 303

  • Boloria (Clossiana) alberta; Pelham, 2008, J. Res. Lepid. 40: 303; [P&N23]
  • Boloria alberta; [BNA]: 322, #246, pl. 26; [Opler]; [MPG] ; [iNaturalist] ; [BOA] ; [BAMONA]
  • Brenthis alberta; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • Clossiana alberta banffensis; [LepIndex]; [CoL]
  • Clossiana alberta; [NACL], #4472; ::NAB, 567; [BARU], #204; [BOW]: pl. 13, f. 17; [BRU2]: 50, pl. 35, f. 1-3; [LepIndex] ; [CoL]

  1600x1200 (~262Kb) ♀ upperside Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.	1600x1200 (~248Kb) ♀ underside Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.	1600x1200 (~233Kb) ♂ upperside Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
  1600x1200 (~226Kb) ♂ underside Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.

  Larva on Dryas octopetala, Vaccinium [BARU], Dryas octopetala, Dryas integrifolia [BRU2]: 50

• Boloria polaris (Boisduval, [1828]) Arctic Europe, Arctic Asia, Arctic America. See [maps] Polarpärlefjäril Polar Fritillary Polaris Fritillary Pohjanhopeatäplä

  • Argynnis polaris Boisduval, [1828]; Eur. Lepid. Index Meth.: 15; TL: Nordkap Peninsula [Norway]
  • Brenthis polaris ab. transversa Lingonblad, 1944; Notulae Ent. 24 (1-2) (1-2): 50; TL: Finland, Malla

  • Boloria (Clossiana) polaris polaris; [P&N23]
  • Boloria (Clossiana) polaris; Pelham, 2008, J. Res. Lepid. 40: 302; [P&N23]
  • Boloria polaris; [BNA]: 323, #248, pl. 26; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34; [MPG] ; [iNaturalist] ; [LEPIFORUM] ; [MBE] ; [MBE] ; [BOA] ; [BAMONA] ; [SVF]
  • Brenthis polaris; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • Clossiana polaris; [NACL], #4470; [H&R], 107; [BOW]: pl. 2, f. 1; [BARU], #202; [BRU2]: 48, pl. 33, f. 1-3; [RFEL]; [LepIndex] ; [CoL]

  660x498 (~61Kb) ♀ upperside FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela

  Larva on Cassiope tetragona, Vaccinium uliginosum, Dryas octopetala, D. punctata [BRU2]: 48

  • B. p. polaris

    • ?Argynnis polaris var. homeyeri Wocke, 1875; in Homeyer, Dt. Nordpolarfart 2: [?? 403 not there]; TL: Grondland
      • = ; Pelham, 2008, J. Res. Lepid. 40: 302
    • Argynnis chariclea var. groenlandica [Skinner], 1892; Ent. News 3 (3): 49, pl. 2, f. 9-10; TL: Greenland
      • = ; Pelham, 2008, J. Res. Lepid. 40: 302
    • Argynnis polaris var. americana Strand, 1905; Rep. 2nd Norweg. Arctic Exp. (3): 10
      • = ; Pelham, 2008, J. Res. Lepid. 40: 303
    • Clossiana alberta kurenzovi Wyatt, 1961; Zs. Wiener ent. Ges. 46: 98, pl. 5, f. 1-2; TL: "Berge von Tschukotka", NE.Siberia
      • = ; Pelham, 2008, J. Res. Lepid. 40: 303
    • Clossiana polaris digna Churkin, 2001; Helios 2: 97, pl. 8, f. 4-6; TL: Wrangel Is., Neizvestnaya R.
      • = ; Pelham, 2008, J. Res. Lepid. 40: 303

    • Argynnis polaris v. americana; Strand, 1912, Ent. Zs. 25 (49): 253
    • Boloria erda kurenzovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Boloria polaris digna; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Boloria polaris polaris; Pelham, 2008, J. Res. Lepid. 40: 302; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34; [BOA]
    • Clossiana alberta kurenzovi; [BARU]; [BRU2], 50; [LepIndex]; [CoL]
    • Clossiana erda kurenzovi; [RFEL]
    • Clossiana polaris americana; [LepIndex]; [CoL]
  • B. p. stellata Masters, 1972 Manitoba

    • Boloria polaris stellata Masters, 1972; Ent. Rec. J. Var. 84 (7/8): 177, pl. 9, f. A, B, H, G; TL: Churchill, Manitoba

    • Boloria (Clossiana) polaris stellata; [P&N23]
    • Boloria polaris stellata; Pelham, 2008, J. Res. Lepid. 40: 303; [BOA]
    • Clossiana polaris stellata; [NACL], #4470a; [CoL]
• Boloria astarte (Doubleday, [1848]) Alaska, Washington, Montana. See [maps] Arctic Ridge Fritillary Astarte Fritillary

  • Melitaea astarte Doubleday, [1848]; Gen. diurn. Lep. (2): pl. 53, f. 5; TL: Rock Lake, Alberta

  • Boloria (Clossiana) astarte astarte; [P&N23]
  • Boloria (Clossiana) astarte; Pelham, 2008, J. Res. Lepid. 40: 303; [P&N23]
  • Boloria astarte tschuktokensis; [iNaturalist]
  • Boloria astarte; [BNA]: 322, #247, pl. 26; [BOC], 285; [Opler]; [MPG] ; [iNaturalist] ; [BOA] ; [BAMONA]
  • Brenthis astarte; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
  • Clossiana astarte; [NACL], #4473; [NAB], 568; [BOW]: pl. 13, f. 18; [LepIndex] ; [CoL]

  Larva on Saxifraga bronchialis [NAB]

  • B. a. astarte NE.British Columbia, Alberta

    • Argynnis victoria Edwards, 1891; Can. Ent. 23 (9): 198; TL: Laggan, Alberta
      • = ; Pelham, 2008, J. Res. Lepid. 40: 303

    • Boloria astarte astarte; [BNA], #247; Pelham, 2008, J. Res. Lepid. 40: 303; [iNaturalist] ; [BOA]
  • B. a. distincta (Gibson, 1920) Yukon - British Columbia

    • Brenthis distincta Gibson, 1920; Rep. Can. arctic exped. 1913-18. 3 (1): 25 i, pl. 4, f. 12; TL: Harrington Creek, Yukon Territory, Canada: lat 65°05'

    • Boloria (Clossiana) astarte distincta; [P&N23]
    • Boloria astarte distincta; Pelham, 2008, J. Res. Lepid. 40: 303; [iNaturalist] ; [BOA]
    • Brenthis distincta; [LepIndex] ; [CoL]
    • Clossiana astarte distincta; [NACL], #4473a
    • Clossiana distincta; [LepIndex] ; [CoL]
    • ?Clossiana distincta; [BARU], #206b; [BRU2]: 51, pl. 34, f. 10-18

    Misidentified/misapplied:

    • Boloria astarte distincta; [BNA], #247

    Larva on Saxifraga spinula, (Russia) Salix , (N.America) [BRU2]: 51

• Boloria tritonia (Böber, 1812) -. See [maps]

  • Papilio tritonia Böber, 1812; Mém. Soc. Imp. Nat. Moscou 3: 20, pl. 1, f. 1-2; TL: [Lake Baikal, Irkutsk, Russia]

  • Argynnis tritonia; [LepIndex] ; [CoL]
  • Boloria tritonia; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34; [iNaturalist] ; [LEPIFORUM]
  • Clossiana tritonia; [BARU], #206a; [BRU2]: 51, pl. 34, f. 7-9, 19-21; [RFEL]; [SZMN]

  Larva on Saxifraga bronchialis [BARU]: 108

  • B. t. tritonia Baikal, Transbaikalia

    • Argynnis amphilochus var. elatus Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 329; TL: Mongolia, Kentey
      • = ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34

    • Boloria tritonia tritonia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana elatus; [BARU], #205
    • Clossiana tritonia elatus; [BRU2], 51
    • Clossiana tritonia tritonia; [BRU2], 51

    Larva on (as elatus) Saxifraga bronchialis, Vaccinium uligonosum [BARU]

  • B. t. amphilochus (Ménétriés, 1859) Amur, Ussuri

    • Melitaea amphilochus Ménétriés, 1859; Bull. phys.-math. Acad. Sci. St. Pétersb. 17 (12-14): 213; TL: Amurland

    • Boloria tritonia amphilochus; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana amphilochus; [LepIndex] ; [CoL]
    • Clossiana tritonia amphilochus; [BRU2], 51; [RFEL]
  • B. t. tschukotkensis (Wyatt, 1961) NE.Yakutia, Chukotka

    • Clossiana distincta tschukotkensis Wyatt, 1961; Zs. Wiener ent. Ges. 46: 99; TL: "Berge von Tschukotka", NE.Siberia
    • Clossiana astarte ershovi Korshunov & Gorbunov, 1995; Butts. Asian Russia: 108; TL: Yakutsk
      • = Boloria astarte tschukotkensis; Pelham, 2008, J. Res. Lepid. 40: 304
    • Clossiana astarte suntara Korshunov & Dubatolov, 1998; New descr. specif. Butts. Asiatic Russia: 21; TL: Suntar-Khayata Range, 180km ESE Khandyga, upper Eastern Khandyga River, 233km on the Kandyga to Magadan line, mtns at 1300-1500m, Yakutia
      • = Boloria astarte tschukotkensis; Pelham, 2008, J. Res. Lepid. 40: 304

    • Boloria (Clossiana) astarte tschukotkensis; [P&N23]
    • Boloria astarte tschukotkensis; Pelham, 2008, J. Res. Lepid. 40: 303
    • Boloria tritonia ershovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Boloria tritonia suntara; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Boloria tritonia tschukotkensis; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana distincta ershovi; [BRU2], 51
    • Clossiana distincta tschukotkensis; [BARU]; [BRU2], 51; [LepIndex]; [CoL]
    • Clossiana tritonia suntara; [RFEL]; [SZMN]
    • Clossiana tritonia tschukotkensis; [RFEL]

    Misidentified/misapplied:

    • Boloria astarte tschukotkensis; [BNA], #247
  • B. t. machati (Korshunov, 1987) Polar Urals

    • Clossiana distincta machati Korshunov, 1987; Novye i maloizvetnye vidy fauny Sibiri, Novosibirsk 19: 10

    • Boloria tritonia machati; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana distincta machati; [BARU]; [BRU2], 51
  • B. t. dubatolovi (Korshunov, 1987) Transbaikalia

    • Clossiana distincta dubatolovi Korshunov, 1987; Novye i maloizvetnye vidy fauny Sibiri, Novosibirsk 19: 11

    • Boloria tritonia dubatolovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana distincta dubatolovi; [BARU]; [BRU2], 51
  • B. t. matveevi (Gorbunov & Korshunov, 1995) Altai

    • Clossiana matveevi Gorbunov & Korshunov, 1995; Butts. Asian Russia: 109; TL: Altai, Kurayskiy Mountain rnage, Aktash, 2700m

    • Boloria tritonia matveevi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
    • Clossiana matveevi; [BRU2]: 51, pl. 35, f. 10-11; [SZMN]
    • Clossiana tritonia matveevi; Yakovlev, 2012, Nota lepid. 35 (1): 91
  • B. t. barkalovi (Dubatolov, 2010)

    • Clossiana tritonia barkalovi Dubatolov, 2010; Amurian Zool. J. 2 (2): 187

    • Boloria tritonia barkalovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34