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Celastrina Tutt, 1906

Celastrina Tutt, 1906; Ent. Rec. 18: 131, TS: Papilio argiolus Linnaeus
Cyaniroides Matsumura, 1919; (preocc. de Nicéville, 1890)

= ; [NL4A, 138]

Maslowskia Kurenzov, 1974; , TS: Celastrina filipjevi Riley

= ; [NL4A, 138]

Celastrina [Wahlberg]
Celastrina [BAMONA]
Celastrina (Lycaenidae) ; [BRU2, 143]
Celastrina (Polyommatini) ; [Opler]
Celastrina (Polyommatinae) ; [NL4A, 138]

Misidentified/misapplied:

Cyaniris; auth., not Dalman, 1816
Lycaenopsis; auth., not C & R. Felder, 1865

600x400 (~80Kb) underside India: Subansiri, 5.11.2005, Photo © Kedar Tokekar

600x400 (~65Kb) underside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

Celastrina argiolus (Linnaeus, 1758)

NAF, EU, Siberia, Turkey, C.Asia, Japan. See [About maps] Paatsamasinisiipi Holly Blue Tosteblåvinge Skogsblåvinge Faulbaumfalter Najade Modraszek wieszczek
Papilio argiolus Linnaeus, 1758; Syst. Nat. (Edn 10) 1 : 483, TL: England
Papilio cleobis Sulzer, 1776; Gesch. Ins. nach linn. Syst.: pl. 18, f. 13-14
Papilio marginatus Retzius, 1783; Gen. Spec. Ins.: 30
Papilio acis Fabricius, 1787; Mantissa Insectorum, 2: 73, n. 687
paraleuca (Röber, 1897)
Lycaenopsis argiolus r. britanna Verity, 1919; Ent. Rec. J. Var. 31: 46
Lycaenopsis argiolus f. calidogenita Verity, 1919; Ent. Rec. J. Var. 31: 46
Celastrina argiolus [Wahlberg]
Celastrina argiolus [Kimmo Silvonen]
Celastrina argiolus [NORLEP]
Celastrina argiolus [Mario Maier]
Celastrina argiolus [Simon Coombes]
Celastrina argiolus ; [BARU, #358]; [EBW]; [BOW: pl. 9, f. 13-14]; [BRU2: 143, pl. 63, f. 16-27, 31-36]
Celastrina argiola ; [MRS, 680]
Celastrina argiolus ; [NL4A, #1086]; [Otakar Kudrna]

500x605 (~31Kb) underside Finland, Tampere, Hervanta, 681:33, 12.5.1996, Photo © Tero Piirainen	533x400 (~41Kb) underside male A road at a birch grove edge near the village Mel'nichikha (Shadrikha), Novosibirsk District and Province, West Siberia, Russia. 18th May 1997, Photo © Oleg Kosterin	1000x1186 (~160Kb) underside Russia, Udmurtia, river banks of Kama, near village Doksha east of Izhevsk, 20.8.2004, Photo © Markku Savela
800x600 (~115Kb) underside Russia, Kaluga area, 19.05.2005, Photo © D. Smirnov	701x851 (~121Kb) male Russia: Kaluga area (35°45'E, 54°23'N), 19.5.2005, Photo © D. Smirnov	500x417 (~56Kb) Turunc, SW Turkey 19/05/92, Photo © Simon Coombes

Larva on Ribes, R. nigrum, R. rubrum, R. uva-crispa, Rhamnus frangula, R. cathartica [SPRK], Caragana frutex, Chamaecystis, Lupinus, Vaccinium [BARU], Berberis, Cuscuta engelmannii, C. lehmanniana, (C. Asia) [BRU2, 144] Sophora flavescens, Lespedeza spp., Desmodium oxyphyllum, Vicia sativa, Pueraria lobata, Wisteria floribunda, Cornus controversa, Aralia elata, Euodia meliifolia, Malus pumila, Prunus salicina, Polygonum cuspidatum, Quercus aliena, Staphylea bumalda [MRS]

C. a. argiolus

EU, Caucasus, Armenia, Talysh Mts., Kazakhstan, Atai, Sayan, W.Siberia, C.Siberia Celastrina argiolus argiolus ; [BRU2, 143]

C. a. mauretanica (Rothschild, 1925)

Morocco, Algeria, Tunisia Lycaena argiolus mauretanica Rothschild, 1925; Novit. Zool. 32 : 214 (repl. algirica Oberthür), TL: Algeria
Lycaena argiolus var. algirica Oberthür, 1915; Étud. Lépid. Comp. 10 : 401, pl. 288, f. 2343, TL: Alger (Algeria)
Celastrina argiolus mauretanica ; [BMAT: 35, pl. 11, f. 25-30]

Larva on Arbutus unedo, Rubus sp., Discolores sp., Cornus sp., Rhamnus sp., Colutea sp. [BMAT]

C. a. bieneri Forster, 1941

Yakutia, Transbaikalia, Far East, Kamchatka Celastrina bieneri Forster, 1941; Mitt. münchn. ent. Ges. 31: 614
Celastrina argiolus bieneri ; [BARU, (note)]; [BRU2, 144]

C. a. caphis (Fruhstorfer, 1922)

Yunnan, ...? Lycaenopsis caphis Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 871 (21 January, 1922)
Celastrina argiola caphis ; [MRS, 680]
Celastrina argiolus caphis ; Huang, 2003, Neue Ent. Nachr. 55: 106 (note)

C. a. sikkima (Moore, [1884])

Sikkim - Assam, Karens Hill Hedge Blue
Cyaniris sikkima Moore, [1884]; Proc. zool. Soc. Lond. 1883 (4): 524, pl. 48, f. 11, TL: Sikkim
Lycaenopsis argiolus sikkima ; [BIR, 280]

Larva on Prinsepia utilis [BIR]

?C. a. kollari (Westwood, [1852])

Chitral - Kumaon Hill Hedge Blue
Lycaena coelestina Kollar, 1848; in Hügel, Kaschmir und das Reich der Siek, 4: 423 (preocc. Lycaena coelestina Eversmann, 1843)
Lycaena kollari Westwood, [1852]; Gen. diurn. Lep. (2): 491 (repl. Lycaena coelestina Kollar, 1848)
Lycaenopsis argiolus kollari ; [BIR, 279(key), 280]

Larva on Prinsepia utilis [BIR]

C. a. iynteana (de Nicéville, 1884)

Nepal - Burma, Thailand, Vietnam, Tibet, S.Yunnan Jyntea Hedge Blue
Cyaniris iynteana de Nicéville, [1884]; J. Asiat. Soc. Bengal, (II) 52 (2/4): 69, pl. 9, f. 7, 7a, TL: Sikkim
Cyaniris jynteana Moore, [1884]; Proc. zool. Soc. Lond. 1883 (4): 524, pl. 48, f. 10, TL: Khasi Hills
Cyaniris victoria Swinhoe, 1893; Trans. ent. Soc. Lond. 1893 (3): 293, TL: Shillong, Assam
Lycaenopsis argiolus var. bothrinoides Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2): 446
Lycaenopsis argiolus var. puspargiolus Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2): 446, TL: Assam
Lycaenopsis argiolus var. albocaeruleoides Chapman, ?1909; ?Proc. zool. Soc. Lond. 1909 (2): 404
Cyaniris jynteana[sic] herophilus Fruhstorfer, 1910; Stettin. ent. Ztg. 71 (2): 293, TL: Vietnam
Cyaniris coelestina cition Fruhstorfer, 1910; Stettin. ent. Ztg. 71 (2): 294, TL: Vietnam
Celastrina argiolus iynteana [Yutaka Inayoshi]
Lycaenopsis jynteana ; [BIR, 280]
Celastrina argiolus iynteana ; Huang, 2001; Huang, 2003, Neue Ent. Nachr. 55: 106 (note)

C. a. hypoleuca (Kollar, [1849])

Kopet-Dagh, Tian-Shan, Ghissar-Darvaz, Pamirs-Alai Lycaena hypoleuca Kollar, [1849]; Denkschr. Akad. Wiss. Wien. 1: 52
Celastrina argiolus hypoleuca ; [BRU2, 143]

C. a. ladonides (d'Orza, 1869)

Amur, Ussuri, Sakhalin, Kuriles, Japan Silvery Hedge Blue
Lycaena ladonides d'Orza, 1869; Lep. Jap.: 20
Lycaena levettii Butler, 1883; Ann. Mag. nat. Hist. (5) 11: 111
Celastrina sachalinensis Matsumura, 1925; J. Coll. Agric. Hokkaido imp. Univ., 15: 105 (preocc.), TL: Sakhalin
Cyaniris heringi Kardakov, 1928; Entomol. Mitt. 17 (?4): 272, TL: Primorye
Lycaenopsis manchurica Matsumura, 1939; Bull. biogeogr. Soc. Japan 9 (20): 358, TL: Manchuria
Celastrina ussuriensis Forster, 1941; Mitt. münchn. ent. Ges. 31: 610
Celastrina ladonides [Wahlberg]
?Lycanopsis ladonides ; [BIR, 279 (key), 280 (Murree - Kumaon)]
Celastrina heringi ; [BARU, #361]
Celastrina ladon ladonides ; [BARU, (note)]
Celastrina argiolus ladonides ; [BRU2, 144]
Celastrina ladonides ; [Oleg Kosterin]

802x642 (~54Kb) underside Deockcheom-ri, Gujwa-eup, Jeju-do Island, South Korea. 31st July 2002, Photo © Oleg Kosterin

744x570 (~38Kb) underside The Shibu River (Shibukawa) shingle right bank at Hidaka village (Hidakamura), Kochi Prefecture, Shikoku, Japan, 5th August 2002, Photo © Oleg Kosterin

638x636 (~52Kb) underside The Shibu River (Shibukawa) shingle right bank at Hidaka village (Hidakamura), Kochi Prefecture, Shikoku, Japan, 5th August 2002, Photo © Oleg Kosterin

Larva on Prinsepia utilis [BIR], Lepedeza bicolor, Vicia amoena, (Ussuri) [BRU2, 144]

C. a. sugurui Eliot & Kawazoé, 1983

Celastrina phellodendroni Omelko, 1987

Ussuri. See [About maps] Celastrina phellodendroni Omelko, 1987; Lepid. Far East: 116, TL: [Gornotaezhnoe, Ussuri]
Celastrina phellodendroni ; [BRU2: 144, pl. 63, f. 37-42]

Larva on Phellodendron amurensis [BARU, Omelko]

Celastrina sugitanii (Matsumura, 1919)

Japan, Korea, NE.China. See [About maps] Lycaena[?] sugitanii Matsumura, 1919; Zool. Mag. Tokyo, 31: 173, TL: Honshu

Larva on (flowers of) Aesculus turbinata, Bothrocaryum controversum [BRU2, 145]

C. s. sugitanii

C. s. ainonica Murayama, 1952

Japan

C. s. kyushuensis Shirôzu, 1943

Japan Celastrina sugitanii kyushuensis Shirôzu, 1943; Zephyrus 9: 189

C. s. sachalinensis (Esaki, 1922)

Sakhalin Cyaniris sachalinensis Esaki, 1922; Zephyrus 9: 265
Celastrina ladon r. sachalinensis ; [BARU, (note)]
Celastrina sugitanii sachalinensis ; [BRU2, 145 (?)]

C. s. lenzeni Forster, 1941

C. s. leei Eliot & Kawazoé, 1983

Celastrina ogasawaraensis (Pryer, 1886)

Japan. See [About maps] Lycaena ogasawaraensis Pryer, 1886; Rhop. Nihon 1: 10, pl. 11, f. 13

Celastrina ladon (Cramer, [1780])

NA, CA. See [About maps] Spring Azure
Papilio ladon Cramer, [1780]; , TL: REF#; pl. 270, f. D;N.America
Argus pseudargiolus Boisduval & Leconte, 1833; Hist. Lép. Am. Sept. (11-22): 118 (1835), pl. 36, f. 1-5
Lycaena violacea Edwards, 1866; Proc. ent. Soc. Philad. 6 (4): 201
deutargiolus (Scudder, 1869); (summer f.)
intermedia (Strecker, 1870); (female)
Celastrina obsoletalunulata Tutt, 1908; (summer f.)
?Celastrina neglectamajor Tutt, 1908; (summer f.)
Celastrina ladon [Wahlberg]
Celastrina neglecta-major [BAMONA]
Celastrina ladon [GALEPS]
?Celastrina neglecta-major ;
?Cyaniris pseudargiolus ; [BOW: pl. 19, f. 11]
Cyaniris ladon ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 45
Cyaniris ladon violacea ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 45
Celastrina ladon ; [NACL, #4363]; [BARU, #359]; [Opler]

identification uncertain 600x440 (~43Kb) underside USA: Bridge Creek Trailhead, North Cascades Highway, (48°30'30N 120°43'00W), Chelan Co., WA, 28.7.1999, Photo © Markku Savela

identification uncertain 800x940 (~96Kb) underside USA: Miller Canyon, Sierra Vista, Cochise Co., Arizona, 8.8.2005, Photo © Markku Savela

Larva on Ceanothus, Cimicifuga racemosa [NAB], Malus, Physocarpus, Prunus, Rubus, Spiraea, Lespedeza, Lupinus, Melilotus, Ledum, Arctostaphylos, Vaccinium, Cimicifuga, Lonicera, Viburnum, Ligustrum, Humulus, Alnus, Cornus, Rhus [BARU]

C. l. ladon

Celastrina argiolus ladon ; [BNA, 397]

C. l. argentata (Fletcher, 1903)

Lycaena argentata Fletcher, 1903; Trans. R. Soc. Canad. 9 (4): 213
Celastrina ladon argentata ; [NACL, #4363b]

C. l. nigrescens (Fletcher, 1903)

Lycaena nigrescens Fletcher, 1903; Trans. R. Soc. Canad. 9 (4): 213
maculatasuffusa (Cockle, 1910); (emend.)
Celastrina ladon nigrescens [ALBERTA]
Celastrina ladon nigrescens ; [NACL, #4363c]

C. l. sidara (Clench, 1944)

Colorado Lycaenopsis pseudargiolus sidara Clench, 1944; Jl N.Y. ent. Soc. 52 (3): 273, TL: Manitou, Colorado
Celastrina ladon sidara ; [NACL, #4363d]

C. l. cinerea (Edwards, 1883)

Arizona Lycaena cinerea Edwards, 1883; Papilio 3 (1) : 8, TL: Arizona
Lycaena pseudargiolus var. arizonensis Edwards, 1884; Trans. Am. Ent. Soc. 11: 306 (summer f.), TL: Arizona

= Celastrina argiolus cinerea ; [NL4A, #1086b]

Cyaniris ladon cinerea ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 45
Cyaniris ladon arizonensis ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 45
Celastrina ladon cinerea ; [NACL, #4363e]
Celastrina argiolus cinerea ; [NL4A, #1086b]

Celastrina echo (Edwards, 1864)

California, Oregon. See [About maps] Lycaena echo Edwards, 1864; Proc. ent. Soc. Philad. 2: 506, TL: California
Cyaniris ladon ab. nunenmacheri Strand, 1915; , TL: REF#b: 159;California

= ; [NL4A, #1086a]

Lycaenopsis pseudargiolus bakeri Clench, 1944; Jl N.Y. ent. Soc. 52 (3): 274, TL: Baker, Oregon

= ; [NL4A, #1086a]

Celastrina echo echo [ILMB]
Celastrina echo echo [BOA]
Celastrina echo [Wahlberg]
Celastrina echo [BAMONA]
Celastrina ladon echo ; [NACL, #4363f]
Celastrina argiolus echo ; [NL4A, #1086a]
Celastrina echo ; [Opler]

Celastrina gozora (Boisduval, 1870)

Mexico, Panama, Honduras. See [About maps] Lycaena gozora Boisduval, 1870; Considérations Lépid. Guatemala: 17, TL: Guatemala (?Honduras, ?Mexico)
Celastrina gozora [ILMB]
Celastrina gozora [BOA]
Celastrina gozora [BAMONA]
gazora ; Holland, 1931, (missp.)
Lycaena gozora ; Godman & Salvin, [1887], Biol. centr.-amer., Lep. Rhop. 2: 104; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 722
Cyaniris ladon gozora ; Dyar, 1903, Bull. U.S. natn. Mus., 52: 45
Celastrina ladon gozora ; [NACL, #4363g]
Celastrina argiolus gozora ; [BNA, 397]; [NL4A, #1086c]

Celastrina nigra (Forbes, 1960)

E.USA. See [About maps] Dusky Azure Spring Sooty Sooty Azure
Celastrina ebenina Clench, 1972; Annls Carnegie Mus. 44 (4): (33-44), TL: Coalburgh, West Virginia
nig (Strecker, 1878); (ab., unavailable)
nigra (Edwards, 1887); (ab., unavailable)
Celastrina nigra [Wahlberg]
Celastrina nigra [BAMONA]
Celastrina ebenina ; [NACL, #4364]
Celastrina nigra ; [BNA: 398, pl. 34, f. 393]; [Opler]

Larva on Aruncus dioicus [BNA]

Celastrina fedoseevi Korshunov & Ivonin, 1990

Transbaikalia, Amur. See [About maps] Celastrina fedoseevi Korshunov & Ivonin, 1990; Taxonom. Ins. Helmint.: 72, TL: Amur region, near Zeya
Celastrina fedoseevi [SZMN]
Celastrina fedoseevi ; [BARU, #360]; [BRU2: 145, pl. 63, f. 28-30]

Celastrina gigas (Hemming, 1928)

W.Himalayas. See [About maps] Lycaenopsis gigas Hemming, 1928; Proc. Ent. Soc. Lond. 3: 29
Celastrina gigas ; [MRS, 682]

C. g. fujianensis Huang, 1994

Fujian Celastrina gigas fujianensis Huang, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 771, 682, f. 74, TL: Fujian

Celastrina hersilia (Leech, [1893])

Nepal - Chinal. See [About maps] Cyaniris hersilia Leech, [1893]; Butts China Japan Corea (2): 319 (1893-1894), pl. 31, f. 16
Celastrina hersilia ; [MRS, 681]

C. h. hersilia

C. h. evansi Toxopeus, 1927

Celastrina evansi Toxopeus, 1927; Tijdschr. Ent. 70 (3/4): 247n

C. h. vipia Cantile & Norman, 1960

Celastrina huegelii (Moore, 1882)

Himalayas (Kashmir - Assam), Ceylon. See [About maps] Large Hedge Blue
Cyaniris huegelii Moore, 1882; Proc. zool. Soc. Lond. 1882 (1): 244
Lycaenopsis huegelii ; [BIR, 279 (key), 280]

Larva on Prinsepia utilis [BIR]

C. h. huegelii

?C. h. singalensis

Ceylon Lycaenopsis huegelii singalensis
?Polyommatus singalensis Moore, 1877; Ann. Mag. nat. Hist. (4) 20 (118) : 342
?Lycaenopsis huegelii singalensis ; [BIR, 280 (note)]

C. h. dipora (Moore, 1865)

Himalayas - Karens, Burma, N.Thailand Dusky Blue Cupid
Lycaena dipora Moore, 1865; Proc. zool. Soc. Lond. 1865 (2): 506, pl. 31, f. 8
Everes huegelii dipora [Yutaka Inayoshi]
Everes huegelii dipora ;
Everes dipora ; [BOR, 270 (key)]

Larva on Desmodium parviflorum, Flemingia fruticulosa [BIR]

C. h. oreoides (Evans, 1925)

Celastrina lavendularis (Moore, 1877)

Dun - Assam, Burma, Bengal, S.India, Ceylon, Malaya, Philippines, Celebes. See [About maps] Plain Hedge Blue
Polyommatus lavendularis Moore, 1877; Ann. Mag. nat. Hist. (4) 20 (118) : 341, TL: Ceylon
Lycaenopsis lavendularis ; [BIR, 278 (key), 279]
Celastrina lavendularis ; [BOW: pl. 175, f. 16]; [MRS, 681]

C. l. lavendularis

C. l. limbata (Moore, 1879)

N.India - Burma, Thailand, Laos, Vietnam, Yunnan Polyommatus limbatus Moore, 1879; Proc. zool. Soc. Lond. 1879 (1): 139, TL: India
?limbata (Moore, 1877); [?]34
Cyaniris placida de Nicéville, [1884]; J. Asiat. Soc. Bengal, (II) 52 (2/4): 68, pl. 9, f. 8, TL: Sikkim
Cyaniris placida hegesander Fruhstorfer, 1910; Stettin. ent. Ztg. 71 (2): 292, TL: Tonkin
Celastrina lavendularis limbata [Yutaka Inayoshi]
Celastrina lavendularis limbata ; [MRS, 681]; Huang, 2003, Neue Ent. Nachr. 55: 106 (note)

C. l. lyce (Grose-Smith, 1895)

Sulawesi Cyaniris lyce Grose-Smith, 1895; Novit. Zool. 2 (4): 506, TL: S.Celebew
Cyaniris pellonia Fruhstorfer, 1910; Stettin. ent. Ztg. 71 (2): 301, TL: Sulawesi

= ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 56

Lycaenopsis limbatus lyce ; Fruhstorfer, 1917, Archiv Naturgesch. (A) 82 (1): 32
Celastrina lavendularis lyce ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 56

C. l. himilcon (Fruhstorfer, 1909)

Taiwan Cyaniris himilcon Fruhstorfer, 1909; Ent. Zs. 23 (25): 117
Celastrina lavendularis himilcon ; [MRS, 681]

C. l. hermesianax (Fruhstorfer, 1910)

lavendularis hermesianax (Fruhstorfer, 1910) (Cyaniris); Stettin. ent. Ztg. 71 (2): 300

C. l. gadara (Fruhstorfer, 1910)

C. l. floresiana (Courvoisier, 1912)

C. l. placidina (Fruhstorfer, 1917)

Sumatra lavendularis placidina (Fruhstorfer, 1917) (Lycaenopsis); Archiv Naturgesch. (A) 82 (1): 31

C. l. isabella Corbet, 1937

Celastrina isabella Corbet, 1937; Trans. R. ent. Soc. Lond. 86 (2): 31
Celastrina lavendularis isabella ; [BMP: 243, pl. 35, f. 38-40]

C. l. chloe Eliot & Kawazoé, 1983

C. l. aroana Eliot & Kawazoé, 1983

?C. l. ssp.?

Serang Celastrina limbata ssp.? ; [BAUR, 381]

Celastrina acesina (Bethune-Baker, 1906)

SE.Papua. See [About maps] Cyaniris acesina Bethune-Baker, 1906; Ann. Mag. Nat. Hist. (7) 17 (97): 103, TL: New Guinea

Celastrina philippina (Semper, 1889)

-. See [About maps] Cyaniris philippina Semper, 1889; Reisen Philipp. (4): 168, pl. 32, f. 14-17 (partim), TL: Philippines

C. p. philippina

Cyaniris philippina philippina ; Fruhstorfer, 1910, Stettin. ent. Ztg. 71 (2): 300
Lycaenopsis cardina philippina ; Fruhstorfer, 1917, Archiv Naturgesch. (A) 82 (1): 14
Celastrina philippina philippina ; Hayashi & Iwanaga, 1974, Kontyu, Tokyo 42: 18, f. 12-13; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 56

C. p. nedda (Grose-Smith, 1894)

Moluccas, Celebes, West Irian - Papua Cyaniris nedda Grose-Smith, 1894; Novit. zool. 1 (3): 572
Celastrina philippina nedda ; [BOW: pl. 175, f. 19]; [BAUR, 381 (text)]

C. p. cinctuta (Grose-Smith, 1895)

Bachan, Halmahera, Ternate philippina cinctuta (Grose-Smith, 1895) (Cyaniris); Novit. Zool. 2 (4): 506
Cyaniris cinctuta Grose-Smith, 1895; Novit. Zool. 2 (4): 506
Celastrina philippina cinctuta[?] ; [BAUR, 381]

C. p. phuste (Druce, 1895)

Timor Cyaniris phuste Druce, 1895; Proc. zool. Soc. Lond. 1895 (3): 573, pl. 32, f. 17
Celastrina philippina phuste ; [BAUR, 381 (text)]

C. p. gradeniga (Fruhstorfer, 1910)

Sulawesi Cyaniris gradeniga Fruhstorfer, 1910; Stettin. ent. Ztg. 71 (2): 301, TL: Sulawesi
Lycaenopsis nedda proba Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 875 (15 April 1922), TL: Sulawesi
Lycaenopsis nedda gradeniga ; Fruhstorfer, 1917, Archiv Naturgesch. (A) 82 (1): 35; Fruhstorfer, 1922, in Seitz, Gross-Schmett. Erde 9 : 875 (15 April 1922), pl. 152, e
Lycaenopsis philippina gradenigra[sic] ; Toxopeus, 1926, 369
Celastrina philippina gradeniga ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 56

C. p. labranda (Fruhstorfer, 1917)

Ambon, Serang, Obi philippina labranda (Fruhstorfer, 1917) (Lycaenopsis); Archiv Naturgesch. (A) 82 (1): 37
?Celastrina philippina labandra[?] ; [BAUR, 381]

C. p. vulcanica (Rothschild, 1915)

Manam I. Lycaenopsis vulcanica Rothschild, 1915; Novit. Zool. 22 (3): 388, TL: Vulcan Is.
Celastrina philippina vulcanica ; [BAUR, 381 (text)]

C. p. lychorida (Fruhstorfer, 1922)

Bismarck Archipelago Lycaenopsis lychorida Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9 : 875 (15 April 1922)
lugra Ribbe²
?lugra (Druce, 1895); Proc. zool. Soc. Lond. 1895 (3): 573, pl. 32, f. 5, TL: Mina Balu
Celastrina philippina lychorida ; [BAUR, 381 (text)]

Celastrina filipjevi (Riley, 1934)

Ussuri, NE.China, Korea. See [About maps] Lycaenopsis filipjevi Riley, 1934; Entomologist 67: 85, pl. 1, f. 1-6, TL: Ussuri
Celastrina filipjevi [Wahlberg]
Maslowskia filipjevi ; [BARU, #362]
Celastrina filipjevi ; [BRU2: 145, pl. 63, f. 43-45]

Larva on Princepia chinensis [BARU, Omelko]

C. f. filipjevi

C. f. admirabilis (Sugitani, 1936)

Celastrina filipjevi admirabilis ;

Celastrina oreas (Leech, [1893])

Ussuri, China, Nepal, NE.India (Assam), Burma, Taiwan. See [About maps] Cyaniris oreas Leech, [1893]; Butts China Japan Corea (2): 321 (1893-1894), pl. 31, f. 12, 15, TL: China, Ta-Chien-Lu
Maslowskia oreas [SZMN]
Celastrina huegeli oreas ;
?Celastrina orcas[sic] ; [BOW: pl. 175, f. 25 (text)]
Maslowskia oreas ; [BARU, #363]
Celastrina oreas ; [MRS, 681]

Larva on Princepia chinensis [BARU, Omelko], Eurya strigillosa, E. acuminata, E. groffii, Prinsepia scandens [MRS]

C. o. oreas

C. o. oreana (Swinhoe, 1910)

Celastrina oreas oreana ;

C. o. arisana (Matsumura, 1910)

Taiwan Cyaniris arisana Matsumura, 1910; Ent. Z. Stuttgart 23 (50): 221, TL: Formosa
Celastrina oreas arisana ; [MRS, 681]

C. o. mirificus (Sugitani, 1936)

Korea, Ussuri Lycaenopsis mirificus Sugitani, 1936; Zephyrus 6: 167, TL: Korea
Maslowskia oreas mirificus [SZMN]
Celastrina oreas mirificus ;
Maslowskia oreas mirificus ; [BARU, (note)]

C. o. septentrionis Forster, 1941

C. o. hoenei Forster, 1941

C. o. baileyi Eliot & Kawazoé, 1983

SE.Tibet (Yigong)

C. o. yunnana Eliot & Kawazoé, 1983

NW.Yunnan Celastrina oreas yunnana ; [MRS, 681]

C. o. limingani Huang, 2001

Celastrina oreas limingani Huang, 2001; Neue Entomologische Nachrichten, 51: 72, pl. 2, f. 16, TL: Gamagou, below Yela Pass, S of Bongda, E.Tibet

C. o. lixianica Sugiyama, 2004

24.9.2009 (26)

References:

[ALBERTA]

University of Alberta, E.H. Strickland Entomological Museum; Browse Entomology Collection

[About maps]

Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

[BAMONA]

Paul Opler, Harry Pavulaan, Ray Stanford, Michael Pogue; Butterflies and Moths of North America; Mountain Prairie Information Node

[BARU]

Korshunov, Y. P., Gorbunov, P. Y.; Butterflies of the Asian part of Russia; English translation by Oleg Kosterin; [Dnevnye babochki aziatskoi chasti Rossii. Spravochnik]

[BAUR]

D'Abrera, 1977; Butterflies of the Australian Region (2nd edn)

[BCA]

electronic; BIOLOGIA CENTRALI-AMERICANA

[BHL]

The; Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!

[BIR]

Wynter-Blyth, M. A., 1957; Butterflies of the Indian Region; (1982 Reprint)

[BMAT]

Tennent, John, 1996; The butterflies of Morocco, Algeria and Tunisia; ISBN 0 906802 05 9

[BMP]

Eliot (Ed.), Corbet & Pendlebury, 1992; The Butterflies of The Malay Peninsula; 4th edition; ISBN 983-9681-05-2

[BNA]

Scott, James A., 1986; The Butterflies of North America

[BOA]

Interactive Listing of American Butterflies; Listado Interactivo de la Mariposas Americanas; Butterflies of America

[BOR]

D'Abrera, B., 1982, 1985, 1986; Butterflies of the Oriental Region, Part I-III

[BOW]

Lewis, H. L., 1974; Butterflies of the World; ISBN 0 245 52097 X

[BRU2]

Tuzov (ed.), Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000; Guide to the Butterflies of Russia and adjacent territories; Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae; Volume 2

[D. Smirnov]

Russia, Moscow; e-mail: D. Smirnov

[EBW]

Smart, Paul, 1976; The Illustrated Encyclopedia of the Butterfly World; ISBN 0 600 31381 6

[GALEPS]

Moths and Butterflies of Georgia and the Southeastern United States; James K. Adams; Dept. of Natural Science, Dalton State College, Dalton, GA ; http://www.daltonstate.edu/galeps/index.htm

[ILMB]

Interactive Listing of Mexican Butterflies

[Kedar Tokekar]

India

[Kimmo Silvonen]

Espoo, Finland; Kimmo's Lep Site

[LAMAS-2005]

Gerardo Lamas; Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)

[LAMAS-NEO]

Gerardo Lamas; Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)

[MRS]

Chou Io (Ed.); Monographia Rhopalocerum Sinensium, 1-2

[Mario Maier]

Europäische Schmetterlinge; e-mail: Mario Maier

[NAB]

Pyle, R. M.; National Audubon Society: Field Guide to North American Butterflies; 1981; ISBN 0-394-51914-0

[NACL]

Hodges, Ronald W. (ed.), 1983; Check List of the Lepidoptera of America North of Mexico

[NHM]

Natural History Museum; The Global Lepidoptera Names Index

[NL4A]

Gerardo Lamas, 2004; Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea - Papilionoidea

[NORLEP]

Norges Sommerfugler

[Oleg Kosterin]

Institue of Cytology & Genetics; Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/

[Opler]

Opler & Warren, 2003 (April); Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.

[SPRK]

Seppänen, E. J, 1970; Suomen suurperhostoukkien ravintokasvit, Animalia Fennica 14

[SZMN]

Lepidoptera; Siberian Zoological Museum

[Simon Coombes]

1 Park Street, Plymouth, PL3 4BL, UK; 01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk

[Tero Piirainen]

Tampere, Finland; e-mail: Tero Piirainen

[Wahlberg]

Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group

[Yutaka Inayoshi]

A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM

[²]

This may require parentheses or not. I don't have the necessary information for this taxon.

Some related literature:

Bethune-Baker, 1906

New species of Lycaenidae from British New Guinea Ann. Mag. Nat. Hist. (7) 17 (97): 100-104

Boisduval, 1870

Considerations sur des Lépidoptères envoyés du Guatemala à M. de l'Orza Considérations Lépid. Guatemala: 100pp

Boisduval & Leconte, 1833

Histoire générale et iconographie des lépidoptères et des chenilles de l'Amérique septentrionale Hist. Lép. Am. Sept. (1): 1-16. pl. 1-3 (1829) (2): 17-24, pl. 4-6 (1829) (3/4): 25-40, pl. 7-12 ([1830]) (5/6): 41-56, pl. 13-18 ([1830]) (7/8): 57-80, pl. 19-24 ([1830]) (9/10): 81-100, pl. 25-30 (1833) (11-22): 101-196, pl. 31-65 ([1835]) (23-26): 197-228, pl. 66-78 ([1837])

Cassidy, 1995

On the Lycaenopsis Group of Genera (Lepidoptera, Lycaenidae) of the Sulawesi Region Trans. lepid. Soc. Japan 46 (2): 45-62

Chapman, 1909

A review of the species of the lepidopterous genus Lycaenopsis Feld. (Cyaniris auct. nec Dalm.) on examination of the male ancilliary appendages Proc. zool. Soc. Lond. 1909 (2): 419-476

Clench, 1944

Two new subspecies of Lycaenopsis pseudargiolus Bdv. & LeC. Jl N.Y. ent. Soc. 52 (3): 273-276

Clench, 1972

Celastrina ebenina, a new species of Lycaenidae (Lepidoptera) Annls Carnegie Mus. 44 (4): 33-44

Cockle, 1910

Notes on a few butterflies found at Kaslo and in northern British Columbia Can. Entomol. 42 (6): 203-204

Corbet, 1937

A Revision on the Malayan species of Celastrina Trans. R. ent. Soc. Lond. 86 (2): 19-33, 12 figs

de Nicéville, [1884]

On new and little known Rhopalocera from the Indian Region J. Asiat. Soc. Bengal, (II) 52 (2/4): 65-91, pls. 1, 9, 10

Westwood, [1852]

The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846) 19-132, pl. 4x, 5-25, 28 (3-14) (1847) 133-200, pl. 1x, 26, 27, 29, 31-44 (15-23) (1848) 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849) 243-250 [?] (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850) 327-466, pl. 59, 67-77 (39-50) (1851) 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)

Druce, 1895

A Monograph of the Bornean Lycaenidae Proc. zool. Soc. Lond. 1895 (3): 556-627, pl. 31-34

Dyar, 1903

A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. natn. Mus., 52: xix, 723pp

Eliot & Kawazoé, 1983

Blue butterflies of the Lycaenopsis group. British Museum (Natural History) Butts. Lycaenopsis group: 309 pp., 560 figs

Edwards, 1862

Descriptions of certain species of diurnal Lepidoptera found within the limits of the United States and of British America. (1-3) Proc. Acad. nat. Sci. Philad. : 160-164 (1861) 54-58 (1862) 221-226 (1862)

Edwards, 1864

Descriptions of certain species of diurnal Lepidoptera, found within the limits of the United States and British America. (3) Proc. ent. Soc. Philad. 2: 501-507

Edwards, 1866

Descriptions of certain species of diurnal Lepidoptera found within the limits of the United States and British America. (5) Proc. ent. Soc. Philad. 6 (4): 200-208

Edwards, 1883

Notes on the collection of butterflies made by Mr. H. K. Morrison in Arizona, 1882 Papilio 3 (1) : 1-10

Edwards, 1883

On the polymorphism of Lycaena pseudargiolus Boisd. Papilio 3 (5/6) : 85-97

Edwards, 1884

Revised catalogue of the Diurnal Lepidoptera of America north of Mexico Trans. Am. Ent. Soc. 11: 245-337 (Separate, Philadelphia, 1884 [1885] 95pp)

Evans, 1915

A list of butterflies caught by Capt. F. M. Bailey in S. E. Thibet during 1913 J. Bombay nat. Hist. Soc. 23: 532-546, 1pl, 1map

Forster, 1941

Neue Lycaeniden-Formen aus China. 2 Mitt. münchn. ent. Ges. 31: 593-627

Fruhstorfer, 1909

Lepidopterologisches Pêle-Mêle. [XVI]. Neues aus Formosa Ent. Zs. 23 (25): 116-117 (18 September)

Fruhstorfer, 1910

Neue Cyaniris-Rassen und Übersicht der bekannten Arten Stettin. ent. Ztg. 71 (2): 282-305 (15 March)

Fruhstorfer, 1917

Revision der Lycaenidengattung Lycaenopsis auf Grund morphologischer Vergleiche der Klammerorgane Archiv Naturgesch. (A) 82 (1): 1-42, 2 pls. (January)

Fruhstorfer, 1922

Gattungen (8-19): Parelodina - Niphanda in Seitz, Gross-Schmett. Erde 9 : 856 (14 February 1921) 857-864 (5 January 1922) 865-872 (21 January, 1922) 873-880 (15 April 1922) 881-888 (18 April 1922) 889-896 (16 May 1922) 897-901 (16 March 1923)

Godman & Salvin, [1887]; Godman & Salvin, [1901]

Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782 3: pl. 1-112

Grose-Smith, 1894

An Account of a Collection of Diurnal Lepidoptera from New Guinea (Parts I-III) - made by Mr. W. Doherty at Humboldt Bay, Dutch New Guinea, and in neighbouring islands, in the Museum of the honourable Walter Rothschild at Tring, with descriptions of new species Novit. zool. 1 (2): 331-365 543-551 (3): 571-584

Grose-Smith, 1895

Descriptions of new species of butterflies, captured by Mr. Doherty in the islands of the Eastern Archipelago, and now in the Museum of the Hon. Walter Rothschild at Tring Novit. Zool. 2 (2): 75-85 (4): 505-514

Guppy & Kondla, 2003

The correct type locality of Cyaniris ladon var. quesnellii Cockle, 1910 (Lycaenidae), with designation of a lectotype J. Lep. Soc. 57 (2): 150-152

Hayashi & Iwanaga, 1974

A new species of Celastrina (Lepidoptera: Lycaenidae) from Borneo and Luzon Kontyu, Tokyo 42: 17-23

Huang, 2001

Report of H. Huang's 2000 Expedition to SE. Tibet for Rhopalocera Neue Entomologische Nachrichten, 51: 65-152

Huang, 2003

A list of butterflies collected from Nujiang (Lou Tse Kiang) and Dulongiang, China with descriptions of new species, and revisional notes Neue Ent. Nachr. 55: 3-114

Kardakov, 1928

Zur Kenntnis der Lepidopteren des Ussuri-Gebietes Entomol. Mitt. 17 (?4): 261-273

Kirby, 1837

Fauna Boreali-Americana or the Zoology of the northern parts of British America in Richardson, Fauna Boreal Amer.: 325pp, 8pls

Leech, [1893]

Butterflies from China, Japan, and Corea Butts China Japan Corea (1): 1-297, pl. 1-28 (1892-1894) (2): 297-681, pl. 29-43 (1893-1894)

Linnaeus, 1758

Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338 339-824

Matsumura, 1910

Lycaeniden Japans Ent. Z. Stuttgart 23 (50): 221-222

Matsumura, 1925

An ennumeration of the butterflies and moths from Saghalien, with descriptions of new species and subspecies J. Coll. Agric. Hokkaido imp. Univ., 15: 83-196, pl. 8-11

Matsumura, 1939

The butterflies from Jehol (Nekka) Manchoukuo, collected by Marqis Y. Yamashina Bull. biogeogr. Soc. Japan 9 (20): 343-359, pl. XIII-XIV

Moore, 1865

List of diurnal Lepidoptera collected by Capt. A.M. Lang in the N. W. Himalayas Proc. zool. Soc. Lond. 1865 (2): 486-509, pls. 30-31

Moore, 1877

Descriptions of Ceylon Lepidoptera Ann. Mag. nat. Hist. (4) 20 (118) : 339-348

Moore, 1879

Descriptions of new Asiatic diurnal Lepidoptera Proc. zool. Soc. Lond. 1879 (1): 136-144

Moore, 1882

List of the Lepidoptera collected by the Rev. J.H. Hocking, chiefly in the Kangra Discrict, N.W Hiamalaya; with Descriptions of new Genera and Species Proc. zool. Soc. Lond. 1882 (1): 234-263, pls. 11-12

Moore, [1884]

Descriptions of new Asiatic diurnal Lepidoptera Proc. zool. Soc. Lond. 1883 (4): 521-535, 2pls [48,?]

Oberthür, 1915

Faune des Lépidoptères de la Barbarie, Algerie Étud. Lépid. Comp. 10 : 1-459, pl. 276-277,277bis,278-315

Omelko, 1987

[Celastrina phellodendroni sp. n., a new sibling species of Celastrina argiolus L. (Lepidoptera, Lycaenidae) from the southern Primorie] Lepid. Far East: 116-122 (Russian)

Opler & Krizek, 1984

Butterflies east of the Great Plains, an illustrated natural history Butts E of Great Plains: 294pp

Riley, 1934

A new Palaearctic lycaenid butterfly Entomologist 67: 85-86

Rothschild, 1915

On the Lepidoptera in the Tring Museum sent by Mr. A. S. Meek from the Admiralty Islands, Dampier and Vulcan Islands Novit. Zool. 22 (2): 192-208 (3): 387-402

Rothschild, 1925

Critical List of the collection of Algerian Lepidoptera of the late Captain N. J. E. Holl. Novit. Zool. 32 : 195-229

Scott & Wright, 1998

A new Celastrina from the Front Range of Colorado Papilio (N.S) 9: 1-15

Semper, 1889

Die Schmetterlinge der Philippinischen Inseln. Reisen im Archipel der Philippinen Reisen Philipp. (1): 1-46, pl. 1-8, A (1886) (2): 47-86, pl. 9-16 (1887) (3): 87-134, pl. 17-24 (1888) (4): 135-174, pl. 25-32 (1889) (5): 175-238, pl. 33-38, B (1890) (6): 239-270, pl. 39-46 (1891) (7): 271-380, pl. 47-49 [iv] (1892)

Sulzer, 1776

Dr. Sulzers Abgekürzte Geschichte der Insecten nach dem Linaeischen System Gesch. Ins. nach linn. Syst.: -

Swinhoe, 1893

A List of the Lepidoptera of the Khasi Hills. (1) Trans. ent. Soc. Lond. 1893 (3): 267-390 (?330)

Toxopeus, 1927

Eine Revision der javanischen, zu Lycaenopsis Felder und verwandten Genera gehörigen Arten (mi Einführung von 6 neuen Genera, Beschreibung von 2 neuen Species un 5 neuen Supspecies. [1] Tijdschr. Ent. 70 (3/4): 232-302, pl. 1, 27 text figs.

Wright & Pavulaan, 1999

Celastrina idella (Lycaenidae: Polyommatinae): a new butterfly species from the Atlantic coastal plain Taxonomic Report 1 (9): 1-11

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.